Abstract not available 

The dentitions as well as one complete and several partial skeletons of Equoids from the Eocene lignite beds of the Geiseltal locality are revised. Instead of 13 species distinguished up to now 3 chronoclines with 5 species and 3 separate species are recognized (text. fig. 1). Propalaeotherium hassiacum HAUPT, 1925 is evolving into Propalaeotherium isselanum (CUVIER, 1824) between the levels of the « obere Unterkohle ›› and the « untere Mittelkohle ›› of the Geiseltal section. Propalaeotherium argentonicum GERVAIS, 1849 is shown to be present in the « untere Unterkohle ››, whereas Lophiotherium pygmaeum (DEPERET,1901) occurs in the « obere Mittelkohle ›› and in the « oberes Hauptmittel ››. Plagiolophus cartieri STEHLIN, 1904 appears during the transition from the « Mittelkohle ›› into the « Oberkohle ›› as the earliest true Palaeothere. Therefore the « Oberkohle ›› is already regarded as Upper Eocene. This is corroborated by the occurrence of a phyletic descendant of Propalaeatherium parvulum (Propalaeotherium n.sp.) in the middle and upper "Oberkohle " because this species appears otherwise for the first time at the mammal level of Lissieu. On the other hand Propachynolophus gaudryz (LEMOINE, 1878) described by Matthes (1977) from the « untere Unterkohle ›› turns out te be in fact a Phenacodont. Thus the decisive argument for classifying the « untere Unterkohle ›› as Lower Eocene has to be dropped. Biostratigraphically the « Unterkohle ›› and the «Basishauptrnittel ›› correspond with the lower Middle Eocene (mammal level of Messel), whereas the «unteres Hauptmittel ›› and the « untere Mittelkohle ›› are equivalent to the middle part of the middle Eocene (mammal level of lssel), and the « obere Mittelkohle ›› together with the « oberes Hauptmittel ›› coincide with the upper Middle Eocene (mammal level of Bouxwiller). 

The Late Miocene site of Gritsev (MN 9, Ukraine) has yielded a very rich bat fauna, the remains of which are well preserved. Compared to other Neogene bat assemblages of Europe, the Gritsev bat community is unique in preserving plecotine bats, which are rare from Neogene sites. Some peculiar and new bat species, including a large plecotin Otonycteris, already were described from the Gritsev mammal site. Here we report new records of small plecotin bats from Gritsev, including a new taxon, Barbastella maxima nov. sp. This is the earliest reliable fossil record of this genus and it differs from more recent species of Barbastella in being considerably larger. The evolutionary patterns in the odontology within the tribe Plecotini, supported by biostratigraphical distribution of fossil records of Plecotus are discussed. The morphological peculiarities of the new fossils of plecotine bats from Gritsev are discussed in connection with its possible taxonomical affinity. 

The exploitation of new early Eocene localities in the Paris Basin has resulted in the collecting of  numerous mammalian remains, among which are about 300 isolated teeth representing the rodents. They belong, for the most part, to the paramyid group. Only the latest level of the early Eocene has yielded rodents belonging to the pseudosciurid group. The paramyids, the object of this study, are represented by at least 5 genera and 10 species; they are distributed among 4 clearly dilferentiated subfamilies : Paramyinae Simpson 1945, Pseudoparamyinae Michaux 1964, Ailuraviínae n. subf., Microparamyinae Wood1962.
It results from this study that the principal types of rodents in the middle and late Eocene of Europe are clearly related to the forms described here: Plesiarctomys Bravard 1850 is related to Pseudoparamys Michaux 1964, Ailuravus Rütimeyer1891 to Meldimys n. gen., Gliravus hammeli Thaler 1966 to Microparamys nanus (Theilard1927), and some Masillamys Tobien 1954 to Microparamys russelli Michaux 1964 and to M. sp. 1. Gliravus and Masillamys are the oldest representatives of the Gliridae and the Pseudosciurídae, respectively, families which will progressively replace the Paramyidae in the course of the middle and late Eocene.
Two stages can be recognized in the history of te European Paramyidae fauna during the course of the early Eocene: the older is characterized by the Mutigny fauna, the younger by the Cuis fauna.
The Mutigny fauna appears already rather diversified and does not completely correspond, point by point, to the homologous fauna of North America. A geographic differentiation seems to have been manifested rapidly, unless the fauna established in Europe was already a little different from that which established itself in North America. 

The comparative morphology and biometry of the lower cheek teeth of modern Equus are studied on approximately 300 mandibles belonging to the 10 usually recognised species : Equus grevyi, E. burchelli, E. quagga, E. zebra, E. africanus, E. asinus, E. hemionus, E. klang, E. przewalskii, E. caballus. The studied parameters comprise : occlusal length and width, postflexid length and index ; shape of the double knot (metaconid + metastylid + lingual groove) ; depth of the vestibular groove on the molars ; frequency of the pli caballinid, protostylid and other enamel plications or islets ; frequency of the dP/l.
The same methods of study are applied to a number of North American, Eurasian and African species. For the sake of comparison, some Hemphillian equids were observed (Dinohippus interpolatus, Dinohippus leidyanus, Astrohippus ansae, Phiohippus mexicanus) but most of the discussed material belongs to Pliocene or Pleistocene species of Equus : the « stenonine ›› E. stenonis, E. simplicidens, E. sanmeniensis and E. teilhardi; the « caballine ›› E. scotti, E. lambei, E. Iaurentius, E. mosbachensis, E. germanicus, E. gallicus, E. taubachensis and the Liakhov horse. The relationships of other species, in particular the North American E. calobatus, E. occidentalis, E. cf mexicanus are not clear for the moment. ln Africa, the Plio-Pleistocene species from Koobi Fora (Kenya) show some stenonine and perhaps asinine affinities. The relationships of E. numidicus and E. tabeti are uncertain but these species are probably related to the East African ones. E. mauritanicus is most certainly related to the Quagga group.
The biometrical data are gathered in 32 tables ; 4 photographie plates and 19 figures illustrate the next. The whole is a complement of the previously published studies of the skulls, upper cheek teeth, incisors and metapodials of modern and fossil Equus.

  

Sieving and washing of about 700 kg of sediment from the miocene site at Montpeyroux produced otoliths of 34 teleost species, of which four still occur in the present day fauna or are near to extant species. Among the fossil species, eight are new : Ilisha lerichei, «genus Clupeidarum ›› orbiculatus, Dipulus mediterraneus, Morone cornuta, Chanda nelsoni, Pomadasys steurbauti, «genus Sciaenidarum ›› barthassadensis and Paraplagusia roseni. The fauna found is typical for a tropical or subtropical very littoral, probably even estuarine environment; it was living in a period near the boundary between Lower and Middle Miocene, perhaps somewhat earlier. 

The study of that locality allowed the description of a new Muridae : Paraethomys athmeniae n. sp. It reveals the existence of new rodent for Algeria : first, a Sciuridae, Atlantoxerus cf. rhodius, and second, a Gliridae, Eliomys truci. So, that work shows the presence of the genus Eliomys in North Africa before the middle of Pleistocene. Lastly, Paraethomys cf. anomalus gives an exact datation of that bed. 

Le présent volume est l'aboutissement d'un projet né il y a presque cinq ans. En décembre 1991, l'un d'entre nous (MG) prenait des contacts en vue de proposer un symposium sur les mammifères fossiles, dédié à D.E. Russell, dans le programme du 4e Congrès de la European Society for Evolutionary Biology. Ce congrès, baptisé "Evolution 93", devait se tenir à Montpellier en août 1993. Son Comité d'Organisation, animé par F. Catzeflis, recherchait des organisateurs de symposiums. L'idée fut acceptée avec enthousiasme par le second d'entre nous (PDG), et le titre de notre Symposium fut précisé: " Palaeobiology and Evolution of Early Cenozoic Mammals - A Symposium in Honor of D.E. Russell". Le projet fut formellement accepté par le Comité d'Organisation en avril 1992. 

As a result of the enrichment of eotheriodont material by one of us (P.K.T.), these specimens (essentially Biarmosuchur and Eotitanosuchur) are reexamined and refigured. A reevaluation of their particularities supports the distinction of two families, for which new diagnoses are proposed. This leads us to discuss the affinities of these families, with respect to the sphenacodonts on one hand, and to the South African primitive theriodonts on the other (gorgonopsids and ictidorhinids). This study contains inherent paleogeographic consequences which are considered in conclusion. 

This paper is about the oldest African rodents faunas, from the late Early Eocene, or early Middle Eocene, Glib Zegdou (Algeria) and Chambi (Tunisia) localities. Five species are described and figured, belonging to a new family here created, the Zegdoumyidae.

This family is compared to the Early and Middle Eocene rodents families from Asia, Europe and North America (Chapattimyidae, Yuomyidae, Gliridae, Theridomyidae, lschyromyidae and Sciuravidae), as well as to those known from the Late Eocene African locality Bir El Ater (Anomaluridae and Phiomyidae).

On the one hand, it seems clear that the African endemic Anomaluridae arise from the Zegdoumyidae. On the other hand, the lschyromyidae, or primitive Sciuravidae, may be the most reliable ancestral groups for the Zegdoumyidae. Thus, this new family can be considered as the sister group for the American Sciuravidae on the one hand, and for the European Gliridae on the other hand.

The biogeographical consequences of these phylogenetic hypotheses are discussed. A new phase of communication between Europe and North Africa is inferred, during the Early Eocene. It has been followed by a short period of endemism, allowing the adaptive radiation for the Zegdoumyidae, preceding the immigration of the Phiomyidae, during the Late Eocene, probably from Asian relatives. 

La localité fossilifère du Puech de Montredon, désignée plus communément sous le nom de Montredon, est située sur la commune de Montouliers (Hérault) à quelques 300 mètres de la limite avec le département de l'Aude. Elle a été découverte en 1845 par Narbonne, Directeur des Mines de La Caunette, et de très nombreux restes de vertébrés continentaux y ont été extraits. La plus ancienne mention de ce gisement dans la littérature semble être celle de Lartet (1859) qui signale que "M. Jourdan, de Lyon, a constaté à Montredon, près de Bize (Aude), l'association des restes de Dinotherium avec l'Hipparion". 

The biometric and morphologie variability of the cheek teeth in the end-of-the-phylum species Lophiodon lautricense Noulet studied in this note, reposes on the observation of about 800 teeth. These were revealed to be little variable in absolute dimensions. The considerable morphologie variability in the upper premolars permitted the problem of the molarization process to be taken up. An hypothesis concerning the order of eruption of the cheek teeth is formulated based on an examination of a large number of milk dentitions. In conclusion, it is suggested that reservations be held on the value of dental characters classically used in systematics for the group under consideration. 

Two steps can be distinguished in the history of the first invasion of western and south western Europe by the arvicolines. The first step corresponds to the installation of these rodents with the immigration of Promimonys inxuliferus Kowalski, then of Mimomys stehlini Kormos and of Mimomys gracilis (Kretzoi). The second is characterized by the establishment of a geographic differentiation in the arvicoline fauna between the south of France and Spain, from where are described new species of Mimomys (Mimamys cappettai, Mimomys septimanus, Mimonys medasensis), and the rest of France, where are found only elements already known from central Europe or England (Mimomys polonicus Kowalski, Mimomys pliocaenicus F. Major, Mimomys reidi Hinton, or forms very close to the latter). This geographic differentiation, which is very certainly the consequence of the division of Europe into distinct climatic provinces, one of them being the southern province comprising at least Spain and southern France, could result from a cladogenetic evolution of Mimomys stehlini and Mimomys gracilis after their immigration. The present work is also a contribution to the search for correlations between the diverse micromammal localities of the latest Pliocene (or early Villafranchian) and of the early Quaternary of Europe. 

The Zanclian marls from Papiol provided otoliths belonging to 53 teleost taxa; nine of those are new for the Mediterranean Pliocene. The association reflects a bathymetry between 150 and 350 m, but it is likely that such depths only existed at the initial stage of flooding of the Llobregat bay. A compilation of the available data for the whole Mediterranean realm at Zanclian time provided a list of 163 taxa, of which 105 could be identified at species level. This last group provides the most useful data for evaluating the composition and affinities of the Mediterranean Zanclian fauna, which appears to be significantly different from the Recent Mediterranean fauna. The Zanclian fauna counts 68 % of Recent species: 12,5 % were already represented in the Mediterranean Miocene, 37 % appeared there at the Zanclian, and 18 % are extra-Mediterranean today. This last group is essentially composed by oceanic fishes, living in the warm parts of the Atlantic and Indo-Pacific Oceans. It is evident that a non neglectable number of Recent species only penetrated very recently in the Mediterranean, which stresses the differences between the more oceanic Zanclian Mediterranean fauna and the present one.


  

Tetraceratops insignis is known from a single, crushed skull from the Lower Permian of Texas. Its unique proportions and osteological details gained central meaning in the question of the origins of Therapsida since this early synapsid has been determined as the oldest and less derived therapsid. Apart from Tetraceratops, the ‘mammal-like’ Therapsida and their sister, the pelycosaur-grade Sphenacodontidae, are separated by one of the longest ghost lineages in tetrapod fossil record. However, the minor, though well justified critique faced insistent publication regarding the therapsid hypothesis. A carefull re-evaluation of the holotypic skull reveals that therapsid traits cannot be supported, including a rejection of the formerly supposed adductor shelf in the temporal fenestra. Increased understanding of ‘pelycosaur’ character variation underlines a haptodontine-grade or, less likely, sphenacodontid position for Tetraceratops. 

Fossils of the endangered sturgeons and peddlefishes are widely distributed. We here report for the first time the presence of one of the extinct osteichthyes genus Cylindracanthus (Liedy 1856a) from the Early Eocene lignite-bearing successions of the Kutch Basin, India. The present well preserved rostrum is characterised by numerous wedge-shaped components encircling the central canal that runs along its length, paired at the base and each wedge contributing to the formation of a ridge. The rostrum lacks teeth. The present find extends the palaeobiogeographical distribution of Cylindracanthus considerably and supports its Eocene age as dental remnants preserved in Cylindracanthus sp. shows a decrease in remanent dentition and tooth bases from the Cretaceous to the Eocene. Cylindracanthus is an useful palaeoenvironmental indicator as it has been found associated typically with deposits of nearshore marine environments. 

Pangolins have never shown a high taxic diversity and their fossil record is scarce. We report here the first discovery of a partial humerus from late Oligocene deposits in Pech du Fraysse (MP28, France). The new specimen from Pech du Fraysse is described and compared to various extant and extinct species of pangolins. It shows a suite of morphological features very similar to the humeri discovered in Saint-André (MP 26), Solnhofen (Burdigalien), and Saulcet (Aquitanian), attributed here to Necromanis franconica. The description of the specimen from Pech du Fraysse allowed us to discuss the systematics of Paleogene and Neogene pholidotans. The differences between PFY 4051 and N. franconica on the one side, and N. quercyi on the other side, might be sufficiently important to justify a generic distinction. A comparison with extant species showed that N. franconica was likely terrestrial and fossorial based on its humeral morphology. 

Previous chronological arrangements of South American Quaternary land mammal faunas are appraised on the basis of current geological and paleontological data. Three South American late Pliocene-Pleistocene land mammal ages are conventionally recognized, from oldest to youngest, the Uquian, Ensenadan, and Lujanian ; all are defined on Argentine faunas.

     The Uquian is based fundamentally and historically on the fauna from the Uquía Formation in Jujuy Province, northwestern Argentina. Important known formations in Argentina yielding Uquian Age faunas include the sub-surface Puelche Formation (or Puelchense) near the city of Buenos Aires, and the Barranca de Los Lobos and Vorohué Formations between Mar del Plata and Miramar, Buenos Aires Province. A tentative subdivision is propos-ed for the Uquian into three subages based on knowledge of the Mar del Plata-Miramar sequence, from oldest to youngest, the Barrancalobian, Vorohuean, and Sanandresian. In Argentina the Uquian is presently marked by the first known record of Scelidodon, Hydrochoeropsis, Ctenomys, Canidae, Ursidae, Gomphotheriidae, Equidae, Tapiridae, Camelidae, Cervidae, and the last known record of Thylatheridium, Thylophorops, Dankomys, Eumysops, Pithanotomys, Eucoelophorus, Hegetotheriidae, Sparassocynidae, and Microtragulidae.

The Ensenadan Age is based on the fauna from the Ensenada Formation near the city of Ensenada, Buenos Aires Province. In Argentina the Ensenadan is marked by the first known record of Lomaphorus, Neothoracophorus, Plaxhaplous, Cavia, Lyncodon, Lutra, Galera, Smilodon, Dicotyles, Lama, Vicugna, the last known record of Orthomyctera, and the only known record of Brachynasua.

     Typícal beds of late Lujanian Age in Argentina consist of fluvial deposits occupying stream channels, and shallow basins, often incised into beds of early Lujanian (i.e. Bonaerian of early workers) and Ensenadan Age. The Lujanian Age is based on a fauna from beds along the Rio Luján, about 65 km west of the city of Buenos Aires, Buenos Aires Province. The Lujanian in Argentina is marked by the first record of Equus, Chlamyphorus, and Holochilus, and the last record of Megatherioidea, Glyptodontoidea, Arctodus (=Arctotherium), Smilodon, Litopterna, Notoungulata, Proboscidea, Equidae, Morenelaphus, and Palaeolama.

   These land mammal ages are often difficult to recognize in other South American countries. The compositions of South American Pleistocene faunas vary with the environment. Some taxa were widely distributed in fossil deposits throughout the continent, but their occurrences need not reflect synchroneity. This is a result of changing climates and habitats in time. Consequently, proposed intracontinental correlations need confirmation based on magnetostratigraphy and a radioisotope time scale. Paleontologic characterizations of these land mammal ages (i.e. first and last record, and guide fossils) are useful for much of Argentina, but extensions to most of the other parts of South America are at best tenuous.

The majority of known non-Argentine Pleistocene faunas are believed to be Lujanian in age. Possible non Argentine early Pleistocene (Uquian) faunas include Ayo Ayo and Anzaldo in Bolivia, and Cocha Verde in southern Columbia. A possible middle Pleistocene (Ensenadan or early Lujanian) fauna is the Chichense of Ecuador. Paleomagnetic and radioisotopic date (MacFadden et al., 1983) clearly indicate that the greater part of the Tarija fauna (Bolivia) is Ensenadan in age.

  The end of the Pleistocene and beginning of the Holocene in South America is marked by extinction of nearly all large mammalian herbivores and their specialized large predators. Radiocarbon age determinations suggest that large scale extinctions of megafauna occurred between 15,000 and 8,000 yrs. B.P. (years before present). 

Study of pantolestid mammals from the late Ypresian locality of Prémontré (reference level MP 10) allows the recognition of three genera: Palaeosinopa. Pantolestes and Premontrelestes n. gen. Pantolestes, up to now only recorded from North America, is represented by P. sabatieri n. sp. Premontrelestes n. gen., represented by the type species P. duchaussoisi n. sp., is compared with Pantolestes and Buxolestes. The latter, known from middle Eocene sites in Europe, has not been recognized at Prémontré. Two other taxa, the first of which is close to Pantolestes and the second of Premontrelestes n. gen. are left in open nomenclature.    

The locality of Montredon has provided 8 species of camivores, often little documented, that are discribed and depicted. All the remains are fragmentary and generally badly preserved which lowers very much the possible precision of the taxonomic study; this one has however allowed the creation of a new subspecies. The most richlypdocumented forms are an ursid (Indarctos) and a felid (Machairodus). Issuing often from a westem Europe evolution, the carnivores are well inserted between those of the better known faunas of the "Classical Pontian" and of the lower Vallesian and this situation fits also well with their stratigraphic level, that cannot however be more accurately defined with them alone.