The oldest european marsupials are described from some specimens (isolated upper molars) recently found from the Hainin sediment (Middle Paleocene of Belgium). These fossils document a new species of the Peradectes genus. They give evidence of a much older occurrence of the marsupials in Europe than it was assumed. They allow us to postulate a didelphid dispersal from South America towards the western-holarctic area operating in two phases : the first one of the Peradectes genus at the end of the Cretaceous; the second one of the Didelphíni tribe at the end of the Paleocene. A central american crossing is likely for the first one,  whereas a transafrican way is tentatively argued for the second one. 

A rear skull from the Quercy Phosphorites is described. It documents a new perissodactyl genus, likely assignable to the family Palaeotheriidae and probably paleogene of age. Owing to the lack of any tooth, this family assignment remains however somewhat hypothetical. The specimen is firstly characterised by the presence of a wide epitympanic sinus swelling and hollowing the squamosal shell and the post-glenoid process. This cavity might make up a peculiar pattern of improvement for the hearing apparatus by carrying out a kind of drum near the middle ear, whereas the bony tympanic remains barely bulged and forms but a few developed auditory bulla. This pattern appears an outcome of a trend observed with many palaeotheriids, such as Plagiolophus. Furthermore, the endocranial cast shows a rather high degree of gyrencephaly for a paleogene mammal. The prominent lambdoidal crest points out a powerful nape musculature and a lowered head bearing. Consequently, it is assumed that such an animal was probably living in somewhat open places, at the expense of rather tough vegetables. It might have been accordingly provided with a semi-hypsodont, cement covered, "plagiolophoid" dentition. 

The analysis of oxygene isotope variations as well as paleobotanical data suggest that the Oligocene/Miocene boundary corresponds to a transitional period marked by floristical and climatic variations. During this period, the pyreneo-alpine tectonics has contribued to modify the geography and western Europe landscapes. Faunal changes (appearances, extinctions, migrations) are observed in different mammalian groups, notably in the rodents. A study of the evolutionary trends and patterns in paleogene rodents is involved for the period ranging from level MP 28 of the Late Oligocene to the Early Miocene, including the Oligo-Miocene boundary.
The Rodents fauna from the sites of Venelles (Bouches-du-Rhône District, France) and Thezels (Lot, France), previously mentionned in litterature, have been studied. The first description of the Eomyidae of La Milloque (MP 29) has been completed. These faunas are compared to those from various localities dating from the considered period. In La Milloque, a new representative of the Eomys species is described next to a form close to Rhodanomys hugueneyae ENGESSER, 1987. It is the Eomys milloquensis nov. sp., the likely descendant of Eomys quercyi COMTE & VIANEY-LIAUD, 1987. Two new species are also described in Thezels: Eucricetodon thezelensis nov. sp., resulting from a likely and local evolution of Eucricetodon praecursor (SCHAUB, 1925) from La Milloque, which, in the same geographic area, could be at the origin of Eucricetodon hesperius ENGESSER, 1985 from Paulhiac. Plesiosminthus admyarion nov. sp., quite distinct from Plesiosminthus schaubi VIRET, 1926, which announces Plesiosminthus myarion SCHAUB 1930. Venelles 'Plesiosminthus schaubi population is considered as a sub-species, named Plesiosminthus schaubi meridionalis nov. subsp. New phylogenetic patterns are proposed. Among the Eomyidae, a quantification of various features of the M1-2/ crown (hypsodonty, degree of abrasion, occlusal angle, state of development of the I and V anticlines), and a comparison with the occlusal diagram of the other teeth among various other populations allows a more efficient separation of Eomys and Rhodanomys genera. In Western Europe, and within this period, it finally does not seem possible to gradually connect the genus Eomys to the genus Rhodanomys. The evolution of the Eomys quercyi - milloquensis lineage seems to underline a similar evolution to that which may have led from the Eomys to the Rhodanomys form. The latter which appears totally accomplished at level MP 29 of the Oligocene is considered as an immigrant. If we compare the most representative species of the Venelles, Thezels, and Coderet sites, (i.e. Rhodanomys, Eucricetodon, Adelomyarion, Peridyromys, Plesíosminthus), it becomes impossible to confirm their biochronological separation. The noticeable differences between the populations may be interpreted as geographical variations. An explanation to these variations, and to fauna's evolution during the Late Oligocene and Early Miocene can be found in the environmental modifications, supported by isotopic, paleobotanical and sedimentologic analysis. A tentative reconstruction of the environments is attempted by the cenogram method. The analysis of the fluctuations of fauna's diversity shows variations which may be correlated to a drop in temperature at MP 29, during the Late Oligocene, followed by an increase in temperature along with an aridity phenomenom, during the basal Miocene (MN O).The confrontation of various methods give the opportunity of reconstituting and comparing the evolution of the environment of three sequences of sites chosen from different regions. Ecological affinities of various rodents' species are being examined. It is possible to consider that the integration of all the conclusions resulting from this study should lead to an explanation to the evolution of rodents for the period around the Oligocene-Miocene boundary. The site of Coderet- level 3- would be posterior to the latter, at the beginnig of the Miocene, and would mark the level MN 0 of the Aquitanian.

  

The Pantolestidae from the middle eocene of Bouxwiller are the subject of a detailed study. Buxolestes hammeli (n. g., n. sp.) is not closely related to any other European or North American form described until now; it presents, however, some characters in common with Pantolestes, a form of the same age from North America. A parallel evolution from a common ancestral form could explain this ressemblance.
Another form (gen. and sp. indet.) accompanies Buxolertes hammeli in the Bouxwiller fauna.
  

 A new genus of hystricognath rodent with two new species, Soriamys gaimanensis and Soriamys ganganensis, from the Colhuehuapian Age (Early Miocene), Sarmiento Formation, of Patagonia is described. The first species comes from the south barranca of the valley of Rio Chubut, near Gaiman locality (Chubut Province). It is known through a great number of dental remains in different stages of wear. The teeth are protohypsodont, with cement in the principal valleys. P4 are simpler, more oval inoutline and without a differentiated hypocone-hypoflexus. Upper molars are pentalophodont in early stages of wear and become bilobated in middle and advanced stages; M3 have a temporary third posterior lobe due to the maintenance of the braquiflexus on the posterior wall of the tooth. Lower teeth have a completely asymrnetrical pattern in relation to the uppers. The molars are trilophodont, with only one complex crest in the trigonid composed of the fusion of the anterolophid and mesolophid. The dp4 have a derived pentalophodont pattern due to the loss or fusion of the mesolophid and the development of an accessory transverse crest between the anterolophid and metalophid. The second species is known through a complete skull and jaw and other skull fragments with the dentition, proceeding from Pampa de Gan Gan, Chubut province. Respect to the first species, the teeth are higher crowned, with an earlier occlusal simplification and a third posterior lobe of M3 more developed and persistent. The skull and jaws show many chinchilloid characters, such as jugal bone with an ascending process, concave palate, very prominent dorsal shelf of the masseteric fossa, nasolachrymal duct opened laterally on the maxillary and very developed lateral mandibular fossa. Moreover, Soriamys is closely related to Cephalomys by sharing a similar asymmetric dental pattern and other dental and mandibular characters. These two genera constitute a natural group, the Cephalomyidae, with peculiar dental characters, like the asymmetric pattem of the upper and lower teeth; Cephalomyopsis, and probably Litadontomys, can be considered part of this group. Scotamys and Perimys constitute a distinct lineage (the Neopiblemidae or Perimyidae), more closely related to the chinchillids. Many characters shared by cephalomyids, eocardiids and caviids suggest a closer affinity between the Chinchilloidea and the Cavioiidea as a whole. Abrocoma shows also an asymmetric dental pattern and other chinchilloid features that suggest some degree of relationship with the cephalomyids, still not well determined. 

A partial skull of Ardynomys russelli sp. nov. (Rodentia, Cylindrodontidae) is described. This was collected in the late Eocene of Alag Tsab locality in the eastem Gobi Desert, Mongolia. Ardynomys russelli sp. nov. is characterized by small size, brachyodont molars, and retention of P3. It represents the earliest record of the genus Ardynomys MATTHEW & GRANGER, 1925, in Asia. 

A lot of isolated teeth of a pteropodid fruit bat has been recently found within an assemblage of micromammals recovered from a karstic fissure filling named Lo Fournas-Il near the locality of Baixas (Pyrénées-Orientales, France). The fauna is Middle Miocene Serravallian age. The fossil fruit bat appears morphologically close to Rousettus; its size is that of a recent medium-sized fruit bat. While the fruit bats are very poorly known as fossils, this discovery shows that one of their recent types of dentitions was perfectly established by Middle Miocene times, and supports the presumed long geologic story of the suborder. One of the major invasions of the Old World fruit bats, supposed originated from SE Asia, reached up to Europe. A suborder unit is added to the miocene fauna of this continent. 

The study of three new populations of Plesiosminthuspromyarion from the "phosphorites du Quercy" and of material from "Auvergne" brings new data on european oligocene Dipodidae. They appear in Western Europe at the beginning of late Oligocene. Evolutionary trends of the group are drawn and particularly the emergence of morphotypes announcing P. schaubi, from the Coderet level, is revealed among the most recent populations of P. promyarion. Differences are attempted to be drawn between the three species : P. promyarion, P. myarion and P. schaubi by restudying the type-population of P. myarion from the aquitanian deposits of Chavroches (Allier) in comparison with two other populations from the same age and the same region. Relationships between early european, american and asiatic Dipodidae are discussed. 

The stapes is the last of the middle ear ossicle chain and the smallest bone of the mammalian skeleton. Because it rests on the membrane of the fenestra vestibuli during life, the stapes may often fall within the bony labyrinth cavity when soft structures decay after death. In this work, we highlight the unexpected role that the bony labyrinth plays in the preservation of the stapes. Systematic investigation of the bony labyrinth of 50 petrosal bones of extinct and extant artiodactyls led to the discovery of eight cases of “intralabyrinthine” stapes. Three dimensional reconstructions of these stapes allow documenting stapes morphology of four extinct artiodactyl taxa: Microstonyx erymanthius (Suidae), Elomeryx borbonicus (Hippopotamoidea), ?Helohyus plicodon (Helohyidae), and an undetermined Cainotheriidae; and four extant ones Choeropsis and Hippopotamus (Hippopotamidae), and Tayassu and Phacochoerus (Suoidea). ?Helohyus plicodon from the Middle Eocene documents the oldest stapes known for the order Artiodactyla. Morphological study and metric analyses of our sample of artiodactylan stapes show that stapes are likely to carry relevant phylogenetic characters/signal within artiodactyls, and a potential Euungulata signature. 

Morphological and biometrical description of skulls, teeth, and limb bones of extant and fossil Old World herniones (including E. hydruntinus) and of New World 'stilt-Iegged' and other slender species from Blancan to Holocene. An Appendix presents ways in which the approximate size of some missing bones or dimensions may be deduced from available ones.

The discussed and/or illustrated fossils were found in Bolivia (Tarija), Canada (Yukon), China (Choukoutien, Gulongshan, Jiling, Loufangzi), Ecuador (Oil Fields), Ethiopia (Melka Kunturé), France (Lunel-Viel), Germany (Süssenborn), Greece (Agios Georgios, Petralona), Hungary (Dorog), Italy (Romanelli), Mexico (Cedazo, San Josecito), Mongolia (Sjara-osso-gol), Spain (Venta Micena), ex-Soviet Union (Akhalkalaki, Binagady, Chokurcha, Chukochya, Kabazi, Kolyma, Krestovka, Kurtak, Staroselie, Tologoj), USA (Alaska, Arkalon, Cedar Meadow, Channing, Conkling, Dry Mountains, Hay Springs, Leisey Shell Pit A, Lissie Formation, Natural Trap, Pool Branch, Powers Ranch, Rock Creek, San Diego, Santo Domingo, Seymour Formation, Shelter, Slaton, Trinity River). Numerous raw or statistically elaborated data are given in Tables.

There is no evidence for the existence of Old World hemiones in the New World nor of 'stilt-Iegged' equids in the Old World. The first 'stilt-Iegged' equid was found at Santo Domingo, New Mexico, and is believed to be Late Blancan. It was probably at the origin of E. calobatus (Arkalon, Rock Creek) and of the smaller E. semiplicatus (Channing, Rock Creek). Slender, but not 'stilt-Iegged', equids found at Natural Trap, Wyoming, ca. 12 ky ago belong to Amerhippus. AlI these species share with Oid World Sussemiones (and some hemiones) peculiar patterns on the lower cheek teeth.

The slender Equus sp. B of Leisey Pit A, Florida, ca. 1.2 Ma, as weIl as Amerhippus francisci and E. tau (probably a senior synonym of E. quinni) share conventional lower cheek teeth patterns. The skulls of A. francisci and E. tau, however, are quite different.

Paleontological data suggest a common origin of Amerhippus, Sussemiones, and 'stilt-Iegged' equids during the late Blancan. Old World hemiones seem to have differentiated later. 

A recently discovered, almost complete specimen of a hybodont tooth, allows us to describe as a new species, fairly common, but usually fragmentary teeth in the Navesink Formation of New Jersey: Hybodus novojerseyensis nov. sp.
Three additional taxa are also described : Heterodontus creamridgensis, Squalicorax sp. and Pseudocorax affinis which is noted for the first time in North America.
The whole elasmobranch fauna of the Early Maastrichtian localities of New Jersey is reevaluated and several generic assignments are changed in comparison to the list previously published in 1975 (Cappetta
& Case). 

Two new Squalicorax species, S. benguerirensis nov. sp. and S. microserratus nov. sp. are described from the Lower Maastrichtian of the Benguérir phosphate open mine, Ganntour deposit, Morocco. The species S. benguerirensis nov. sp. was classically assigned to S. yangaensis since Arambourg (1952) and has been also recognized in coeval deposits from eastern USA to Mid-East. The species S. microserratus nov. sp. correspond to the lateral teeth of S. kaupi as reported by Arambourg (1952) and which is now referred in fact to S. bassanii. The comparison of these two new species with other Anacoracids, known in Moroccan or elsewhere, allows highlighting the great taxonomic and ecological diversities of this family during the Cretaceous.
  

A new middle Miocene hipposiderid bat is described from a limestone deposit on Riversleigh Station in north-western Queensland. Hipposideros (Brachipposideros) nooraleebus n. sp. is the first record of this subgenus from anywhere in the world outside of France. The palaeoecological setting of the fossil bats appears to have been a relatively quiet, sunny lime-enriched tropical pool that contained tortoises, crocodiles and fish. It is possible that the bats were washed into the pool from an adjacent cave.
The Riversleigh bat most closely resembles the French Burdigalian (early middle Miocene) bat H. (B.) aguilari. It is also possible that it may have been closely related to the original Australian hipposiderid stock that ultimately gave rise to the endemic monotÿpic Rhinonycteris aurantius. The disjunct distribution of species of H. (Brachipposideros) suggests that representatives of this subgenus will be found in at least tropical southern Asia. 

The composition of assemblages of lizards and Amphisbaenian from the European Eocene are described. At least ten lizard families are identified from the lower European Eocene levels. Eight are still recorded in the last level (Escamps) of the late Eocene. Agamid lizards (genus Tinosaurus) died out by the end of the lower Eocene and Varanid lizards (genus Saniwa) disappeared by the beginning of the late Eocene. Amphisbaenians are recorded throughout the Eocene in Europe. The lacertilian fossil record of Europe and North America show a high degree of faunal resemblance in the early Eocene, followed by a decrease during the later part of the epoch. The lacertilian and amphisbaenian faunas from the European Eocene are not subject to great variations during the period; this is in contrast with the mammal record at the same time. It is argued that the low metabolic rates and the ectothermy of lizards could explain those differences, along with the increasing insularity of the West European area during the late Eocene time.

  

Study of pantolestid mammals from the late Ypresian locality of Prémontré (reference level MP 10) allows the recognition of three genera: Palaeosinopa. Pantolestes and Premontrelestes n. gen. Pantolestes, up to now only recorded from North America, is represented by P. sabatieri n. sp. Premontrelestes n. gen., represented by the type species P. duchaussoisi n. sp., is compared with Pantolestes and Buxolestes. The latter, known from middle Eocene sites in Europe, has not been recognized at Prémontré. Two other taxa, the first of which is close to Pantolestes and the second of Premontrelestes n. gen. are left in open nomenclature.    

La faune de Rongeurs d'Escamps (Lot) bien que relativement pauvre en espèces (dix) s'avère riche d'enseignement pour les Rongeurs de l'Oligocène inférieur d'Europe Occidentale. Cette periode semble caracterisée par une cladogenèse des Théridomyines. A Escamps, un nouveau genre (Patriotheridomys) est décrit ainsi qu'une nouvelle espèce de Theridomys. Avec Oltmamys platyceps, décrit ici plus complètement et désormais bien situé stratigraphiquement, les deux espèces précédentes constituent un ensemble original du Sud de la France. A la même époque, en Angleterre, Allemagne et Espagne, des espèces «régionales» de Théridomyinés se dersifient.  A côté de ces formes qui ne semblent pas franchir la «Grande Coupure» sont représentées deux lignées d'Issiodoromyinés (Elfomys sp et Pseudoltmomys cuvieri), une de Théridomyiné (T. (Blainvillimys) rotundidens) ainsi qu'un Gliridé, Gliravus priscus (que l'on différencie nettement de la deuxième lignée des Gliravus oligocènes : G.  meridionalis -> G. majori). On retrouve ces formes dans les gisements plus récents du niveau de Hoogbutsel où elles sont à peine plus évoluées. Le fait que les degrés évolutifs de ces lignées soient très proches laisse supposer que le laps de temps séparant les niveaux d'Ecamps et Hoogbutsel fut relativement court. 

In the course of the current revision of adapid primates from the Eocene Geiseltal, an almost complete specimen was found in the Geiseltal Museum collections. The fossil, the most complete adapid specimen so far discovered in Europe, has been determined as Pronycticebus neglectus n. sp.
Ecology and locomotion as well as the likely phylogenetic position within the infraorder Adapiformes are discussed. 

The skull remains referred to the genus Palaeotherium are recorded and described. Biometrical tests are made to elucidate intrageneric allometric relationships and to allow comparisons with various other perissodactyls. Apart from the well known shortness of post canine diastems and deepness of the narial opening, the genus is characterized by a great lengthening of the splanchnocranium, owing to a spreading of the post-orbital facial region, by a reduced area of the eye-socket and by the prevalence of the temporal muscle with regard to the masseter; this original shape of the masticatory apparatus needs to be related to the morphology of the jugal teeth and particularly to their  asymmetrical semi-hypsodonty.
    These animals, whose running ability was evidently poor, appear to have been adapted to rather closed environments, feeding on relatively soft vegetable matter; olfactory sense was likely to play a leading part in interindividual and environmental relationships. Such evolutionary trends might explain the disappearance of most of them, as clirnatic conditions deteriorated at the end of the Eocene, before the "Grande Coupure" which affected mammalian faunas at that time.
      Although the present paper is not directly concemed with phylogenetics, it invalidates the supposed ancestor-descendant relationship between P. castrense and P. magnum, and it suggests a possible emergence of the P. medium lineage from a P. siderolithicum stock. Moreover, the structure of the post-orbital facial area allows the  erection of a new sub-genus, Franzenitherium, for the  species lautricense and duvali.

  

Les Carnivores Fissipèdes de Sainte-Néboule appartiennent tous au genre Cynodictis et semblent constituer une population homogène. Celle-ci se distingue suffisamment des espèces déjà décrites pour constituer un taxon particulier : Cynodictis lacustris neboulensis n. s. sp. . L'étude des variations à l'intérieur de cette population nous a conduit à reconsidérer les critères utilisés pour définir les espèces existantes et à regrouper certaines d'entre elles. Il semble qu'il demeure cependant trois lignées distinctes dans le genre Cynodictis mais le matériel nous paraît encore insuffisant pour traduire cette remarque en termes de systématique. 

The petrosal of the Paulchoffatiidae HAHN, 1969 is described and compared with that of younger multituberculates and of other Mesozoic mammals. The "Morrison petrosal", described by Prothero (1983), is also discussed; it probably belongs to the multituberculates. The reconstruction of the ventral side of the Paulchoffatiinae-skull, given by Hahn in 1987, is completed by addition of the otic and the occipital region.