Abstract not available 

The new fauna of Collet Redon (Bouches-du-Rhône, France) is dated by three rodents: Megacricetodon aff. bavaricus, Democricetodon affinis mutilus and Peridyromys cf. hamadryas. They correlate this locality with Oggenhof and Ohningen in Bavaria (Western Germany). As the radiometric age of Ohningen is estimated between 14 and 13 M.Y., these three localities are of Serravallian age. This datation brings a complete readjusment of the stratigraphy of the section of Collet Redon formerly described by Collot and Combaluzier. The marine deposits with underly the continental formation with the mammal fauna, are Burdigalian. The angular unconformity between the marine and the continental deposits gives evidence of an episode of emersion on the margin of a sedimentary basin, with deformation and erosion. Owing to the newly discovered fauna, this geodynamical event is clearly settled within the regional geographical and chronological context. Lacustrine and continental deposits of such an age were up to now unsuspected in this area. 

Two new Squalicorax species, S. benguerirensis nov. sp. and S. microserratus nov. sp. are described from the Lower Maastrichtian of the Benguérir phosphate open mine, Ganntour deposit, Morocco. The species S. benguerirensis nov. sp. was classically assigned to S. yangaensis since Arambourg (1952) and has been also recognized in coeval deposits from eastern USA to Mid-East. The species S. microserratus nov. sp. correspond to the lateral teeth of S. kaupi as reported by Arambourg (1952) and which is now referred in fact to S. bassanii. The comparison of these two new species with other Anacoracids, known in Moroccan or elsewhere, allows highlighting the great taxonomic and ecological diversities of this family during the Cretaceous.
  

The composition of assemblages of lizards and Amphisbaenian from the European Eocene are described. At least ten lizard families are identified from the lower European Eocene levels. Eight are still recorded in the last level (Escamps) of the late Eocene. Agamid lizards (genus Tinosaurus) died out by the end of the lower Eocene and Varanid lizards (genus Saniwa) disappeared by the beginning of the late Eocene. Amphisbaenians are recorded throughout the Eocene in Europe. The lacertilian fossil record of Europe and North America show a high degree of faunal resemblance in the early Eocene, followed by a decrease during the later part of the epoch. The lacertilian and amphisbaenian faunas from the European Eocene are not subject to great variations during the period; this is in contrast with the mammal record at the same time. It is argued that the low metabolic rates and the ectothermy of lizards could explain those differences, along with the increasing insularity of the West European area during the late Eocene time.

  

The present study of Artiodactyla from the Middle Eocene of the Geiseltal lignite beds concems systematics, biostratigraphy, and palaeoecology on the basis of 174 specimens: isolated remains to more complete skeletons. Instead of the formerly known five species of two families are now recognized 14 species of the Diacodexeidae, Dichobunidae, Cebochoeridae, and Haplobunodontidae. New species are Aumelasia maniai, Anthracobunodon neumarkensis, Masillabune franzeni. Four species of the Geiseltalfauna are definitely known from elswere, and five species are closely related to those from other European localities. Evidently the faunal situation of artiodactyls during the Middle Eocene of Europe was largely uniform. The distribution of artiodactyls within the sequence of the Geiseltal strata corroborates the biostratigraphical concept of the land mammal age Geiseltalian (Franzen & Haubold l986b) as well as the mammalian reference levels MP 11-13 (Franzen 1987). Reconstructions of the skulls and skeletons allow conclusions on the functional morphology and palaeoecology of the artiodactyls of the European Middle Eocene 

Le matériel provenant de nouvelles fouilles dans les phosphorites du Quercy, soumis aux divers spécialistes, a conduit à la soutenance de plusieurs thèses d'état: sur les rongeurs (Hartenberger, Vianey-Liaud), les Chiroptères et Insectivores (Sigé), les Artiodactyles (Sudre), les Squamates (Rage) et, en partie, les Chéloniens (De Broin). Chacun dans son domaine, à côté des conclusions d'ordre évolutif, paléogéographique ou paléoécologique, a pu établir des successions stratigraphiques des gisements du Quercy qui se sont révélées largement concordantes. Ainsi la succession des faunes du Quercy est actuellement l'une des mieux établies. Dans ce contexte, les différents spécialistes ont décidé de conjuguer leurs efforts dans l'étude monographique de plusieurs gisements repérés le long de cette échelle, afin de rassembler l'information paléontologique sur des faunes bien précises et de tenter d'obtenir des indications d'ordre taphonomique. 

An unpublished mandible of the large hyracoid Titanohyrax andrewsi from the early Oligocene Jebel Qatrani Formation, Fayum Depression, Egypt is described. This specimen has a twofold importance. Firstly, it opens an unexpected window on early paleontological research in the Fayum because it was discovered as early as 1904 by the French paleontologist René Fourtau during an expedition to the Fayum organized by the Muséum National d’Histoire Naturelle, Paris (MNHN). This expedition has remarkably never been mentioned in the literature. Secondly, the mandible documents the best-preserved specimen of T. andrewsi, permitting a revision of one of the very rare Paleogene hyracoids. Interestingly, the new mandible was discovered two years before the first report of the species by Charles W. Andrews. The hypodigm of T. andrewsi is reviewed and the dentition as a whole is compared in detail, notably with other Titanohyrax species from the Fayum. The validity of the large Titanohyrax “schlosseri” species is discussed, but a pronounced sexual size dimorphism for T. andrewsi is favoured. 

HGL50 is a latest Early-early Middle Eocene vertebrate-bearing locality located in Western Algeria. It has produced the richest and most diverse fauna of amphibians and squamate reptiles reported from the Palaeogene of Africa. Moreover, it is one of the rare faunas including amphibians and squamates known from the period of isolation of Africa. The assemblage comprises 17 to 20 taxa (one gymnophionan, one probable caudate, three to six anurans, seven ‘lizards’, and five snakes). Two new taxa were recovered: the anuran Rocekophryne ornata gen. et sp. nov. and the snake Afrotortrix draaensis gen. et sp. nov. The locality has also yielded the first confirmed anilioid snake, the first Palaeogene gymnophionan, and probably the first caudate from the Palaeogene (and possibly from the Tertiary) of Africa. The presence of a caudate at that time in Africa would be of particular interest; unfortunately, the available material does not permit a definitive identification. The fauna comprises Gondwanan and more specifically West Gondwanan vicariants, probably autochthonous groups and a Eurasian immigrant (assuming that the identification of the caudate is accurate). The fauna from HGL50 is clearly distinguished from the few other Eocene assemblages of Africa. However, if this results largely from differences in geological ages, geographic positions of the localities and mainly differences in environments took a part in the composition of the faunas. 

Eosimiid and amphipithecid primates document a long and significant history of primate evolution throughout the Eocene in Southeast Asia. Despite the absence of a comprehensive post-Eocene fossil record, it was generally hypothesized that both families left no descendant in Asia. Recently, two new small-bodied taxa, Bugtipithecus and Phileosimias, have been recovered in early Oligocene coastal deposits from the Bugti Hills (Balochistan, central Pakistan) and referred to the families Amphipithecidae and Eosimiidae, respectively, on the basis of dental fossil remains. In this paper, we provide more exhaustive description, comparison, and discussion of these taxa. As for tarsiid and sivaladapid primates, the persistence of eosimiids and amphipithecids into the Oligocene clearly demonstrates that low latitudes of South Asia provided a continuous access to tropical refugia during the climatic deterioration characterizing the late Eocene-early Oligocene interval, which was seemingly lethal for primate communities elsewhere across the Holarctic continents. As a contribution to the ongoing phylogenetic debates regarding the position of eosimiids and amphipithecids on the primate family tree, we have performed a cladistic analysis in a high-level primate systematic context in order to assess the position and the role of these new taxa in that phylogenetic issue. Our results support the view according to which eosimiids and amphipithecids (and by extension Phileosimias and Bugtipithecus, respectively) are stem anthropoids. These fossils from Pakistan document an unsuspected Oligocene phase of the evolutionary history of anthropoid primates in southern Asia, which clearly enhances the extent of the anthropoid radiation in this province during the Paleogene. Several phylogenetic and paleobiogeographic aspects are discussed, notably the intra- and inter-relationships between Paleogene Asian and Afro-Arabian anthropoids, and the resulting potential dispersal models between both land-masses during the Paleogene. 

The Las Águilas site near Porvenir de Jalpa, Coahuila, Mexico, is extremely rich in tetrapod remains comprising both bones and trackways of several dinosaur taxa of late Campanian age. Within a 50 m thick section we identified at least nine layers with dinosaur bone assemblages. In one of these the dinosaur bones are associated with remnants of eusuchian crocodilians, turtles, plesiosaurs, pterosaurs, tyrannosaurids, dromaeosaurids, parksosaurid, hadrosaurids, ceratopsids, and ankylosaurs. This layer is also rich in coprolites of turtles, crocodilians and likely theropods, thus providing evidence for the wealth of Late Cretaceous vertebrate life in the area. 

Until now Aplodontidae of the European Oligocene have been documented by four species only. The phylogenetic relations remained obscure. as the distribution of only one species has been known in some detail. New material made it possible to define the stratigraphic range of two of the already existing species (Plesispermophilus angustidens, Sciurodon cadurcense) and to follow their development during the Oligocene beginning with the event of the « Grande Coupure ››. Sciurodon remained nearly without change until the end of the Middle Oligocene. Plesispermophilus angustidens split into two distinct phyletic lines, one of which (P. macrodon n. sp.) reaching considerable size, is represented till the beginning of the Upper Oligocene (Pech de Fraysse, Gaimersheim). The other line leads to Plesispermophilus ernii (basal Upper Oligocene of Burgmagerbein 1. terminal Upper Oligocene of Coderet). Besides the already known forms a new small-sized species (P. atavus n. sp.) is described, which by its primitive features closely resembles the genus Plesispermophilus. Two other small-sized species already known from the Upper Oligocene (? P. argoviensis) and Lower Miocene (? P. descedens) seem to be closely related to the new species. It cannot be decided whether they are descendents of this line or have developed independently, because of their poor fossil record.
Comparison of the evolutionary modalities in the different phylogenetic lines reveals general trends. the most striking of which is the complication of the tooth pattern by the development of additional crests. In the lower molars the cusps diminuate in size and are more and more transformed into ridges. ln addition new connection between the crests appear. in the upper molars, the « selenodont » shape of the teeth becomes more and more dominant, and in the two main evolutionary lines of Plesispermophilus the metaconulus becomes duplicated. A further evolutionary trend is the size increase of the premolars compared to the molars, which is even more pronounced in the Miocene Aplodontidae.
Phylogenetic relations between the primitive Plesispermophilus and certain « prociurines ›› of Northern America as well as between Plesispermophilus (P. angustidens) and more progressive forms of the Upper Oligocene (P. ernii, P. macrodon n. sp.) can be documented. In this light, the taxonomic distinction between Prosciurinae (bunodont) and Allomyinae (selenodont) sensu Rensberger 1976 can be shown to be artificial, because it separates forms from each other, which are evidently closely related. Consequently the separation into two subfamilies has been abolished. 

The ventral face of the Paulchoffatiinae skull (Multituberculata, Lower Kimmeridgian, Portugal) is new reconstructed. Some details hitherto unknown are added, as the presence of jugals, the structure of the palatine and the extension of the pterygoids. The situation of the m2/ is discussed. Kielanodon hopsoni n. g., n. sp. is erected, known by its upper p3-5/. From Guimarotodon leiriensis the mandible with its dentition is made known. New informations concerning the milk-dentition and the replacement of teeth are also added. 

The Middle Eocene Fossillägerstätte of the Geiseltal lignite beds near Halle/S. (German Democratic Republic) is the reference locality of the European land mammal age Geiseltalian and of the Mammalian Paleogene reference levels MP 11 - MP 13. Due to this importance a reinvestigation is given of the lithostratigraphical development of the Geiseltal beds and of their vertebrate sites. The last are genetically related to the southwest border of the Geiseltal depression and the influx of carbonate-rich waters. The geographical distribution and stratigraphical position of the fossiliferous sites depends on subrosive and tectonically controlled distribution of coal seams. The geological factors and the known stratigraphical guide of some mammalian species suggest corrections of the age of some sites. Four of the alltogether five coal bearing phases contain the 35 sites with mammalian remains. By the distribution of the around 69 mammal species are characterized, with 5 faunal steps ranging from MP 11 to MP 14 or over the Geiseltalian up to the Lower Robiacian. Well distant are the faunas of MP 11 and MP 12. Beginning with MP 12 up to MP 13/14, the fossil record is very frequent by 27 sites. This evidence coincides somewhat more with the concept of land mammal ages compared to that of the punctual mammalian reference levels. 

The study of another faunal assemblage (mostly macromammals) from Torralba de Ribota (Calatayud, Zara-
goza Prov.) demonstrates the Middle "Burdigalian" age of the deposit, MEIN zone 4a. Some ten genera have been recognized. a.o. Anchitherium, Aceratherium and Lagopsis. 

Sainte-Néboule has yielded only 4 species of Rodents. But the Theridomyids (Blainvillimys rotundidens and Patriotheridomys altus) are very significative of the age of the locality: Ste-Néboule is lower than the marker level of Escamps 

The stapes is the last of the middle ear ossicle chain and the smallest bone of the mammalian skeleton. Because it rests on the membrane of the fenestra vestibuli during life, the stapes may often fall within the bony labyrinth cavity when soft structures decay after death. In this work, we highlight the unexpected role that the bony labyrinth plays in the preservation of the stapes. Systematic investigation of the bony labyrinth of 50 petrosal bones of extinct and extant artiodactyls led to the discovery of eight cases of “intralabyrinthine” stapes. Three dimensional reconstructions of these stapes allow documenting stapes morphology of four extinct artiodactyl taxa: Microstonyx erymanthius (Suidae), Elomeryx borbonicus (Hippopotamoidea), ?Helohyus plicodon (Helohyidae), and an undetermined Cainotheriidae; and four extant ones Choeropsis and Hippopotamus (Hippopotamidae), and Tayassu and Phacochoerus (Suoidea). ?Helohyus plicodon from the Middle Eocene documents the oldest stapes known for the order Artiodactyla. Morphological study and metric analyses of our sample of artiodactylan stapes show that stapes are likely to carry relevant phylogenetic characters/signal within artiodactyls, and a potential Euungulata signature. 

A general view is given of the vertebrate faunas, Cretaceous to Quaternary of age, found in Columbia and of their principal characteristics. This view leads to the discussion of the isolation of the South American continent and of the role played by the Bolivar syncline with respect to North American immigrants during the Oligocene. The absence of marine deposits of Oligocene age in the north and northwest of Columbia suggests the possibility of a communication with Central America. This communication would have permitted the passage of hystricomorph rodents, of platyrrhine monkeys, and of colubrids. The non-occupation, until then, of the ecologie niches of these groups would have favored their installation beside the indigenous fauna. In this hypothesis it would no longer be necessary to admit that these vertebrates arrived as «island hoppers ››. The eco-biologic conditions would explain the absence of large-sized forms of North American origin. 

Two steps can be distinguished in the history of the first invasion of western and south western Europe by the arvicolines. The first step corresponds to the installation of these rodents with the immigration of Promimonys inxuliferus Kowalski, then of Mimomys stehlini Kormos and of Mimomys gracilis (Kretzoi). The second is characterized by the establishment of a geographic differentiation in the arvicoline fauna between the south of France and Spain, from where are described new species of Mimomys (Mimamys cappettai, Mimomys septimanus, Mimonys medasensis), and the rest of France, where are found only elements already known from central Europe or England (Mimomys polonicus Kowalski, Mimomys pliocaenicus F. Major, Mimomys reidi Hinton, or forms very close to the latter). This geographic differentiation, which is very certainly the consequence of the division of Europe into distinct climatic provinces, one of them being the southern province comprising at least Spain and southern France, could result from a cladogenetic evolution of Mimomys stehlini and Mimomys gracilis after their immigration. The present work is also a contribution to the search for correlations between the diverse micromammal localities of the latest Pliocene (or early Villafranchian) and of the early Quaternary of Europe. 

Afin d'étudier la magnétostratigraphie de la colline de Montredon, j'ai prélevé en Avril 1982, 93 carottes orientées réparties sur 29 sites. J.-P. Aguilar et I. Loridant m'ont aidé durant cette opération. Pour la stratigraphie de la coupe échantillonnée et pour le contenu faunique des deux gisements de mammifères de cette colline, voir J.-P. Aguilar & J.-Y. Crochet (1982).
[...] 

About the early radiation of Rodents, we have now from the early tertiary of Asia, a new fossil record, and
we can do new interpretations. First the problem of the origin of Rodents is studied : considered as a sister group of Lagomorpha, it is possible to find their ancestors between the Mixodontia. Second the new facts about the origin of modern groups of Rodents are reviewed. A general scheme of this diversification can be proposed.