A lot of isolated teeth of a pteropodid fruit bat has been recently found within an assemblage of micromammals recovered from a karstic fissure filling named Lo Fournas-Il near the locality of Baixas (Pyrénées-Orientales, France). The fauna is Middle Miocene Serravallian age. The fossil fruit bat appears morphologically close to Rousettus; its size is that of a recent medium-sized fruit bat. While the fruit bats are very poorly known as fossils, this discovery shows that one of their recent types of dentitions was perfectly established by Middle Miocene times, and supports the presumed long geologic story of the suborder. One of the major invasions of the Old World fruit bats, supposed originated from SE Asia, reached up to Europe. A suborder unit is added to the miocene fauna of this continent. 

Pangolins have never shown a high taxic diversity and their fossil record is scarce. We report here the first discovery of a partial humerus from late Oligocene deposits in Pech du Fraysse (MP28, France). The new specimen from Pech du Fraysse is described and compared to various extant and extinct species of pangolins. It shows a suite of morphological features very similar to the humeri discovered in Saint-André (MP 26), Solnhofen (Burdigalien), and Saulcet (Aquitanian), attributed here to Necromanis franconica. The description of the specimen from Pech du Fraysse allowed us to discuss the systematics of Paleogene and Neogene pholidotans. The differences between PFY 4051 and N. franconica on the one side, and N. quercyi on the other side, might be sufficiently important to justify a generic distinction. A comparison with extant species showed that N. franconica was likely terrestrial and fossorial based on its humeral morphology. 

Two new species, Omanodon minor n. g., n. sp. and Shizarodon dhofarensis n. g., n. sp., known from fifteen isolated teeth, are described here as the first adapiform primates (?Anchomomyini) recognizable to date in the Taqah material (early Oligocene of Sultanate of Oman).

Omanodon minor n. g., n. sp. displays special morphological similarity to the adapid tribe Anchomomyini from the Eocene of Europe, and especially to the Anchomomys lineage. Resemblances with the extant lemurifonn Microcebus are also noticeable and could be regarded as supporting Schwartz & Tattersall (1983) hypothesis of special relationships between the anchomomyine adapids and the cheirogaleid lemuriformes. However, these morphological affmities can be interpreted, altematively, as the results of parallelisms: important differences in upper molars indicate that the resemblances of cheirogaleids and Omanodon minor n. g., n. sp. are indeed probably due to parallelisms. Phyletic relationship of O. minor n. g., n. sp. to Anchomomyini is finally the most likely hypothesis.

Shizarodon dhofarensis n. g., n. sp., although much more poorly known, is closely related to Omanodon minor n. g., n. sp., at least at a familial level. The general morphology of this species suggests
also a close link with adapid Anchomomyini, although precise relationships within this tribe remain obscure. Interesting resemblances of Shizarodon dhofarensis n. g., n. sp. to Djebelemur martinezi lower molars (early Eocene of Tunisia) are also noticeable. These resemblances are even stronger than those betwen Omanodon minor and Djebelemur martinezi. However the very bunodont upper molars referred to D. martinezi are unusual for adapids, and there are moreover some notable differences in their lower molars. Thus resemblances in Djebelemur and Shizarodon are probably due to paralellisms.

Because of the fragmentary nature of the material and of possible parallelisms, the systematic position of Omanodon and Shizarodon within adapiformes cannot however yet be established definitively. 

The dentitions as well as one complete and several partial skeletons of Equoids from the Eocene lignite beds of the Geiseltal locality are revised. Instead of 13 species distinguished up to now 3 chronoclines with 5 species and 3 separate species are recognized (text. fig. 1). Propalaeotherium hassiacum HAUPT, 1925 is evolving into Propalaeotherium isselanum (CUVIER, 1824) between the levels of the « obere Unterkohle ›› and the « untere Mittelkohle ›› of the Geiseltal section. Propalaeotherium argentonicum GERVAIS, 1849 is shown to be present in the « untere Unterkohle ››, whereas Lophiotherium pygmaeum (DEPERET,1901) occurs in the « obere Mittelkohle ›› and in the « oberes Hauptmittel ››. Plagiolophus cartieri STEHLIN, 1904 appears during the transition from the « Mittelkohle ›› into the « Oberkohle ›› as the earliest true Palaeothere. Therefore the « Oberkohle ›› is already regarded as Upper Eocene. This is corroborated by the occurrence of a phyletic descendant of Propalaeatherium parvulum (Propalaeotherium n.sp.) in the middle and upper "Oberkohle " because this species appears otherwise for the first time at the mammal level of Lissieu. On the other hand Propachynolophus gaudryz (LEMOINE, 1878) described by Matthes (1977) from the « untere Unterkohle ›› turns out te be in fact a Phenacodont. Thus the decisive argument for classifying the « untere Unterkohle ›› as Lower Eocene has to be dropped. Biostratigraphically the « Unterkohle ›› and the «Basishauptrnittel ›› correspond with the lower Middle Eocene (mammal level of Messel), whereas the «unteres Hauptmittel ›› and the « untere Mittelkohle ›› are equivalent to the middle part of the middle Eocene (mammal level of lssel), and the « obere Mittelkohle ›› together with the « oberes Hauptmittel ›› coincide with the upper Middle Eocene (mammal level of Bouxwiller). 

Abundant crocodilian fossil remains have been collected, mainly by R. Lavocat. from several levels of the
Palaeogene of the Tilemsi valley (eastern Mali). Several dyrosaurids occur in the Palaeocene (Montian ?) : Phosphatosaurus sp., Hyposaurus bequaerti (Dollo). H. wilsoni (Swinton). H. nopesai (Swinton), Rhabdognathus rarus Swinton, R. compressus n. sp. ln the lower Eocene has been found a dyrosaurid which cannot be accurutely determined.In the middle Eocene were found a dyrosaurid, Tilemsisuchus lavocati n. g., n. sp., and an indeterminate eusuchian. 

Some fragmentary remains of spines and endoskeletal elements, referred here to as ctenacanthid like elasmobranchs, are recorded in the Middle Devonian of Bolivia. These specimens, and some others from the Eodevonian of Brasil represent the only  Devonian fish remains hitherto known from South America and indicate that further discoveries may be made in their original localities respectively.
  

The new fauna of Collet Redon (Bouches-du-Rhône, France) is dated by three rodents: Megacricetodon aff. bavaricus, Democricetodon affinis mutilus and Peridyromys cf. hamadryas. They correlate this locality with Oggenhof and Ohningen in Bavaria (Western Germany). As the radiometric age of Ohningen is estimated between 14 and 13 M.Y., these three localities are of Serravallian age. This datation brings a complete readjusment of the stratigraphy of the section of Collet Redon formerly described by Collot and Combaluzier. The marine deposits with underly the continental formation with the mammal fauna, are Burdigalian. The angular unconformity between the marine and the continental deposits gives evidence of an episode of emersion on the margin of a sedimentary basin, with deformation and erosion. Owing to the newly discovered fauna, this geodynamical event is clearly settled within the regional geographical and chronological context. Lacustrine and continental deposits of such an age were up to now unsuspected in this area. 

The Las Águilas site near Porvenir de Jalpa, Coahuila, Mexico, is extremely rich in tetrapod remains comprising both bones and trackways of several dinosaur taxa of late Campanian age. Within a 50 m thick section we identified at least nine layers with dinosaur bone assemblages. In one of these the dinosaur bones are associated with remnants of eusuchian crocodilians, turtles, plesiosaurs, pterosaurs, tyrannosaurids, dromaeosaurids, parksosaurid, hadrosaurids, ceratopsids, and ankylosaurs. This layer is also rich in coprolites of turtles, crocodilians and likely theropods, thus providing evidence for the wealth of Late Cretaceous vertebrate life in the area. 

Near "la Lieude", in the Lodève basin, more than a thousand of footprints are distributed in a twenty of trackways which amounts to 220 m length. They have been found on calcareous siltstone level in the B site named also "Réserve Naturelle Volontaire". This last is located in the Saxonian summit dated Upper Permian. "La Lieude" tracks are described by using statistical methods then they are compared with others from the world Permian. What allows to distinguish 4 following ichnotaxa: Lunaepes ollierorum nov.ichnosp., Merifontichnus thalerius nov. ichnogen. and nov. ichnosp., Planipes brachydactylus nov.ichnosp. and Brontopus circagiganteus nov. ichnosp. All these traces are attributed with possibility or probability to Therapsida or to Therosauria, except Brontopus circagiganteus nov. ichnosp. that could be due to Caseamorpha. All these animals whose sizes have been estimated between l and 5 m lived probably in a playa environment.The biological and sedimentological data from "la Lieude" footprints levels compared with informations provided by the tracks orientations, suggest the following scenario. Animals coming from the North have crossed a sandy channel bank with plants zones by directing to the South for the majority. Maybe, they were going to the lacustrine part of the playa, close to "la Lieude" footprints that they have just trampled on. 

The recently discovered, very rich, Pliocene locality of Layna (Soria, Spain), has already yielded 30 species of mammals. Hipparion fissurae, described here is more dolichopodic than other Hipparion. It is related to Hipparion crusafonti from Villaroya (Villafranchian), but more slender between other characters.
  

The genus Pachynolophus, one of the poorest known of the Palaeotheriidae, includes the brachyodont forms with reduced and non-molariform premolars and with upper molars lacking a mesostyle. Quantitative characters (divers surface indications and elongation of the teeth), while demonstrating a close relationship to Hyracotherium, permit a better differentiation of the genus, confirm its specific splitting, and permit the distinction of three lineages. The skull from Memerlein is taken as the type of a new species, P. Iavocati, of which the dentition is extremely characterized by its lophiodonty, the strong reduction of the premolars and the reduction of the cingula. This characterization testifies to a late age which extends the existence of the genus quite near to the Eocene-Oligocene limit. Compared with the only two skulls known of related species (Hyracotheríum vulpiceps and Pachynolophus Iivinierensis), that from Memerlein is distinguished by progressive characters affecting diferent regions but most particularly the braincase; it is not possible, however, to isolate within this evolution the part which leads to a systematic differentiation. Modernization is translated by a considerable increase in size of the braincase, principally in the frontal region, a development of the facial region with anterior displacement of the dental series and a greater specialization of the masticatory apparatus. This evolution parallels the history of the Equidae of the North American early Tertiary, but certain particularities, the form of the alisphenoid, the presence of an anterior frontal foramen, and the structure of the paroccipital apophysis, testifies to the independance of the European forms. 

The first part of this study was devoted to a descriptive analysis of teeth morphologies among the vespertilionine bats. This leads now to a tentative synthesis, providing views on the systematics of the group. The results could be seen according to three distinct but closely related purposes : 1 - the sorting of the genera contents in order to conform the genera units to homogeneous taxa that could represent natural issues of evolutionary lineages ; 2 - the investigation of relationships between extant genera in order to infer the possibilities of common origin ; 3 - according to the preceeding items and to the observed evolutionary trends, a tentative phylogeny, modest and cautious. The contents of many genera are sorted : Leuconoe is removed from subgeneric to generic position, whereas Myotis becomes a subgenus of it ; the myotodont species are cleared away from the Pipistrellus genus ; Glischropus and Scotozous are synonymized within Pipistrellus ; Hypsugo is raised to the generic level ; some species previously ranged within Pipistrellus will form provisionally a collective group, Attalepharca nov. ; the Eptesicus genus is broken up, the excluded species being grouped within Nycterikaupius gen. nov. ; the Nycticeini tribe is defined again after exclusion of Otonycteris , Scotoecus, Scotophilus , and addition of Hesperoptenus ; the species la io and Pipistrellus tasmaniensis are removed to Eptesicus (n.s.) and Pipistrellus dormeri to Scotoecus. Groupings of genera are stated according to the main evolutionary trends of I1/. The relevance of these is often warranted by close morphologic similarities of other teeth. This leads to a recognition of the major evolutionary radiations which occurred in the group. The filiations schematized at the end of the work show the dental relationships observed between the extant genera, and could represent a phylogenic framework. Two major facts are to be underlined : 1- the early divergence of leuconoids ; 2 - the successives crossings to myotodonty from the nyctaloid flow. Fossil data from the literature are punctually and tentatively incorporated within phylogenic sketches. 

Le présent volume est l'aboutissement d'un projet né il y a presque cinq ans. En décembre 1991, l'un d'entre nous (MG) prenait des contacts en vue de proposer un symposium sur les mammifères fossiles, dédié à D.E. Russell, dans le programme du 4e Congrès de la European Society for Evolutionary Biology. Ce congrès, baptisé "Evolution 93", devait se tenir à Montpellier en août 1993. Son Comité d'Organisation, animé par F. Catzeflis, recherchait des organisateurs de symposiums. L'idée fut acceptée avec enthousiasme par le second d'entre nous (PDG), et le titre de notre Symposium fut précisé: " Palaeobiology and Evolution of Early Cenozoic Mammals - A Symposium in Honor of D.E. Russell". Le projet fut formellement accepté par le Comité d'Organisation en avril 1992. 

Abstract not available 

The discovery in the Turolian (Late Miocene) of Dytiko 3 (Macedonia, Greece) of a complete skull with mandibles and cervical vertebrae, atlas and epistropheus, is a very important contribution to the knowledge of the subfarnily Chalicotheríinae. After the description, the comparison with other specimens of Miocene chalicotheres permits the revival of the generic name Macrotherium with a new species M. macedonicum. This genus is mainly characterized by a short snout and an inflated cerebral skull. It coexists during the Miocene with Chalicotherium. A cladistic analysis leads to conclusion that the species which has been described from the Early Middle Miocene of Rusinga must be identified as the type-species of a new genus: Butleria.





  

A strong lophodonty, an extreme heterodonty, some hypsodonty and regular overlayings of coronal cement are prominent features of the genus Leptolophus (Palaeotheriinae = Palaeotheriidae s.s.). The histological pattern of the teeth unusually joins type II enamel prisms, characteristic of advanced ungulates, together with archaic features, such as an almost complete lack of Hunter-Schreger zonation and a weak expanse of peritubular dentine. The skull is narrow and slender, with an elongated ante-orbital facial region, a moderately notched nasal aperture, a rather elongated post-canine diastem, parallel zygornatic arches and a fairly dorsally located squamoso-mandibular joint.The functional analysis brings to light "ectolophodont" masticatory cycles with two phases, in which maximum power was applied, contrary to equíds, on hindmost teeth; likewise, skull accomodations to increasing height of the teeth are quite different. This study leads to the assumption that Leptolophus may have been light mammals, living in rather open surroundings, browsing on herbaceous plants or leaves cropped close to the ground. Moreover, it appears that it could have been some inadequacy of dental structures to the dietary, which leaded to quick wear of the teeth and to many enamel notches, but had been somewhat balanced by the early increase of hypsodonty, not induced in such a case by a biotop deterioration (as it will happen at the end of the Eocene). This ínadaptation might account for the short duration of the genus Leptolophus, whose the 3 species, L. stehlini, L. nouletí and L. magnus n. sp. are indeed confined in the level MP 16. Its geographical spreading (as far as known, South of western Europe) and the morphological pattern of its dentition suggest that this genus would have been related to early upper Eocene endemic spanish forms.





  

A well-preserved anterior cranium of Paranourosorex gigas RZEBIK-KOWALSKA, 1975 from Podlesice, Poland, provides additional knowledge of this rare shrew, including almost complete maxillary dentition. The systematic status of this species as a relative of recent Anourosorex MILNE-
EDWARDS, 1870 is confirmed 

Eosimiid and amphipithecid primates document a long and significant history of primate evolution throughout the Eocene in Southeast Asia. Despite the absence of a comprehensive post-Eocene fossil record, it was generally hypothesized that both families left no descendant in Asia. Recently, two new small-bodied taxa, Bugtipithecus and Phileosimias, have been recovered in early Oligocene coastal deposits from the Bugti Hills (Balochistan, central Pakistan) and referred to the families Amphipithecidae and Eosimiidae, respectively, on the basis of dental fossil remains. In this paper, we provide more exhaustive description, comparison, and discussion of these taxa. As for tarsiid and sivaladapid primates, the persistence of eosimiids and amphipithecids into the Oligocene clearly demonstrates that low latitudes of South Asia provided a continuous access to tropical refugia during the climatic deterioration characterizing the late Eocene-early Oligocene interval, which was seemingly lethal for primate communities elsewhere across the Holarctic continents. As a contribution to the ongoing phylogenetic debates regarding the position of eosimiids and amphipithecids on the primate family tree, we have performed a cladistic analysis in a high-level primate systematic context in order to assess the position and the role of these new taxa in that phylogenetic issue. Our results support the view according to which eosimiids and amphipithecids (and by extension Phileosimias and Bugtipithecus, respectively) are stem anthropoids. These fossils from Pakistan document an unsuspected Oligocene phase of the evolutionary history of anthropoid primates in southern Asia, which clearly enhances the extent of the anthropoid radiation in this province during the Paleogene. Several phylogenetic and paleobiogeographic aspects are discussed, notably the intra- and inter-relationships between Paleogene Asian and Afro-Arabian anthropoids, and the resulting potential dispersal models between both land-masses during the Paleogene. 

Messiphoca mauretanica nov. gen., nov. sp. représente le premier Phocidae fossile recensé en Afrique du Nord. Provenant du gisement sahélien (Miocène supérieur) de Raz-el-Aïn (Algérie), il est connu par quelques os du membre antérieur (humérus, radius, ulna), quelques vertèbres dorsales et un crâne très fragmentaire. La description de cette forme nouvelle amène à considérer Messiphoca mauretanica comme un Monachinae archaïque proche de l'origine du groupe Pliophoca - Monachus. L'incidence de cette interprétation d'un Phocidae «pré-Messinien ›› sur la crise de salinité du Messinien est aussi envisagée. 

The Condylarths from Hainin (Hainault, Belgium) show no affinity at the generic level to those known in other Paleocene localities of Europe and North America ; they are described as new forms : Monshyus praevius n. gen., n. sp. and Prolatidens waudruae n. gen., n. sp. Monshyus praevius, discovered in only one of the levels in the excavation at Hainin, is similar to the genera Microhyus TEILHARD and Louisina RUSSELL ; with them it is included in the subfamily Louisininae (Hyopsodontidae). With respect to Microhyus and Louisina, Monshyus is distinguished by the precociously modern aspect of its upper molars, the only teeth that are referable. Prolatidens waudruae, known only by lower molars, was found in several levels in the pit at Hainin. It is an arctocyonid presenting possible relationships to the North American form Oxyprimus galadrielae ; it therefore has been provisionally attributed to the subfamily Oxyclaeninae. If this attribution is confirmed, this species will constitute the first and only representative of the group in Europe.