Two new species, Omanodon minor n. g., n. sp. and Shizarodon dhofarensis n. g., n. sp., known from fifteen isolated teeth, are described here as the first adapiform primates (?Anchomomyini) recognizable to date in the Taqah material (early Oligocene of Sultanate of Oman).

Omanodon minor n. g., n. sp. displays special morphological similarity to the adapid tribe Anchomomyini from the Eocene of Europe, and especially to the Anchomomys lineage. Resemblances with the extant lemurifonn Microcebus are also noticeable and could be regarded as supporting Schwartz & Tattersall (1983) hypothesis of special relationships between the anchomomyine adapids and the cheirogaleid lemuriformes. However, these morphological affmities can be interpreted, altematively, as the results of parallelisms: important differences in upper molars indicate that the resemblances of cheirogaleids and Omanodon minor n. g., n. sp. are indeed probably due to parallelisms. Phyletic relationship of O. minor n. g., n. sp. to Anchomomyini is finally the most likely hypothesis.

Shizarodon dhofarensis n. g., n. sp., although much more poorly known, is closely related to Omanodon minor n. g., n. sp., at least at a familial level. The general morphology of this species suggests
also a close link with adapid Anchomomyini, although precise relationships within this tribe remain obscure. Interesting resemblances of Shizarodon dhofarensis n. g., n. sp. to Djebelemur martinezi lower molars (early Eocene of Tunisia) are also noticeable. These resemblances are even stronger than those betwen Omanodon minor and Djebelemur martinezi. However the very bunodont upper molars referred to D. martinezi are unusual for adapids, and there are moreover some notable differences in their lower molars. Thus resemblances in Djebelemur and Shizarodon are probably due to paralellisms.

Because of the fragmentary nature of the material and of possible parallelisms, the systematic position of Omanodon and Shizarodon within adapiformes cannot however yet be established definitively. 

The discovery in the Turolian (Late Miocene) of Dytiko 3 (Macedonia, Greece) of a complete skull with mandibles and cervical vertebrae, atlas and epistropheus, is a very important contribution to the knowledge of the subfarnily Chalicotheríinae. After the description, the comparison with other specimens of Miocene chalicotheres permits the revival of the generic name Macrotherium with a new species M. macedonicum. This genus is mainly characterized by a short snout and an inflated cerebral skull. It coexists during the Miocene with Chalicotherium. A cladistic analysis leads to conclusion that the species which has been described from the Early Middle Miocene of Rusinga must be identified as the type-species of a new genus: Butleria.





  

Rodents from the new localities of Châteauredon (Alpes-de-Haute-Provence) and La Denise (Bouches-du-Rhône) show more similarities with the rodents known from the Czecho-Slovak locality of Franzenbad than with the species found in contemporaneous Lower Miocene faunas located westward of the present Rhône Valley. This is another data wich support the existance of several biogeographical provinces in Southern France during Lower Miocene. The correlations which have been settled down allow new datations of several formations in Provence and give new information on the Aquitanian and Burdigalian paleogeographies of this region. More precise stratigraphical allocations are given to some other localities of Central Europe. 

Messiphoca mauretanica nov. gen., nov. sp. représente le premier Phocidae fossile recensé en Afrique du Nord. Provenant du gisement sahélien (Miocène supérieur) de Raz-el-Aïn (Algérie), il est connu par quelques os du membre antérieur (humérus, radius, ulna), quelques vertèbres dorsales et un crâne très fragmentaire. La description de cette forme nouvelle amène à considérer Messiphoca mauretanica comme un Monachinae archaïque proche de l'origine du groupe Pliophoca - Monachus. L'incidence de cette interprétation d'un Phocidae «pré-Messinien ›› sur la crise de salinité du Messinien est aussi envisagée. 

The intensive exploration of the Pliocene Hadar Formation, rich in hominid remains, led us to the discovery of several micromammals levels. ln some of them, rodents are very abundant. The stratigraphic repartition of these levels do not cover the whole fossiliferous series of the formation but takes place only in the sedimentary members from Sidi Hakoma and Denen-Dora (rancing from 3.1 - 3.2 MY to 2.8 - 2.9 MY, according to the recent geochronological data). During this gap of time, the species do not show morphological changes, what allowed us to gather, in the same taxa, forms of slighty different ages.
Two striking facts, giving a lot of indications, characterize these small rodents'faunas. First, we notice the domination of the Muridae, as well on a qualitative way (number of species) as on a quantitative one (number of individuals). Then, it appears that, until now, two genera of these murids were known only in the south-western asiatic regions. So, we can suppose continuous biotops between Africa and Indian Subcontinent before 3 MY. In this hypothesis, the hominids had already the possibility to leave their african « cradle ››. Finally, almost all studied genera are still represented at the present time. This fact, previously observed in Laetolil, Omo, Olduvai contributes to remove hope of establishing a biochronological scale based on rodents, in tropical zone. Nethertheless, that allows to try a reconstruction of the palaeoenvironnement, by using the principle of actualism.
  

A new species, Otonycteris rummeli nov. sp., is described from the Late Miocene site Gritsev (MN 9) in the Ukraine. Otonycteris rummeli nov. sp. differs from those of most vespertilionids, except recent Otonycteris, Antrozous and Early Miocene Karstala silva, in having a well-developed entocingulid at the foot of the trigonid valley in the lower molars. The morphological resemblance of Otonycteris, Antrozous and Karstala is apparently a case of convergence in the evolution of the Old and New Worlds bat faunas. From at least the Middle Miocene the range of Otonycteris distribution spread to the whole of Central Europe and such a situation continued during the whole Late Miocene. This indicates a more arid climate in Europe during the Upper Miocene compared to the Quaternary. The reduction of the distribution range of Otonycteris and its extinction in most of the territory of Europe could have been caused by the global climatic cooling and increasing glacial cycle amplitude during the onset of the Quaternary. 

The faunal list of the mammals collected at the locality of Aumelas (Hérault, France) is revised. For the first
time this Middle Eocene locality is precisely settled in the european chronological scale of "niveaux repères", between the levels of Bouxwiller and Egerkingen, in Uppermost Lutetian.
A new Dichobunid from the site is described : Aumelasia gabineaudi n. g., n. sp. This new genus has primitive characters. and it may be in the descent of the Lower Eocene Protodichobune. 

The Las Águilas site near Porvenir de Jalpa, Coahuila, Mexico, is extremely rich in tetrapod remains comprising both bones and trackways of several dinosaur taxa of late Campanian age. Within a 50 m thick section we identified at least nine layers with dinosaur bone assemblages. In one of these the dinosaur bones are associated with remnants of eusuchian crocodilians, turtles, plesiosaurs, pterosaurs, tyrannosaurids, dromaeosaurids, parksosaurid, hadrosaurids, ceratopsids, and ankylosaurs. This layer is also rich in coprolites of turtles, crocodilians and likely theropods, thus providing evidence for the wealth of Late Cretaceous vertebrate life in the area. 

Pangolins have never shown a high taxic diversity and their fossil record is scarce. We report here the first discovery of a partial humerus from late Oligocene deposits in Pech du Fraysse (MP28, France). The new specimen from Pech du Fraysse is described and compared to various extant and extinct species of pangolins. It shows a suite of morphological features very similar to the humeri discovered in Saint-André (MP 26), Solnhofen (Burdigalien), and Saulcet (Aquitanian), attributed here to Necromanis franconica. The description of the specimen from Pech du Fraysse allowed us to discuss the systematics of Paleogene and Neogene pholidotans. The differences between PFY 4051 and N. franconica on the one side, and N. quercyi on the other side, might be sufficiently important to justify a generic distinction. A comparison with extant species showed that N. franconica was likely terrestrial and fossorial based on its humeral morphology. 

A fossil species of the extant genus Leuconoe, L. lavocati n. sp. from Le Garouillas Oligocene locality, Quercy phosphorites, SW-France, is established in nomenclatural standards. 

The recently discovered, very rich, Pliocene locality of Layna (Soria, Spain), has already yielded 30 species of mammals. Hipparion fissurae, described here is more dolichopodic than other Hipparion. It is related to Hipparion crusafonti from Villaroya (Villafranchian), but more slender between other characters.
  

The genus Pachynolophus, one of the poorest known of the Palaeotheriidae, includes the brachyodont forms with reduced and non-molariform premolars and with upper molars lacking a mesostyle. Quantitative characters (divers surface indications and elongation of the teeth), while demonstrating a close relationship to Hyracotherium, permit a better differentiation of the genus, confirm its specific splitting, and permit the distinction of three lineages. The skull from Memerlein is taken as the type of a new species, P. Iavocati, of which the dentition is extremely characterized by its lophiodonty, the strong reduction of the premolars and the reduction of the cingula. This characterization testifies to a late age which extends the existence of the genus quite near to the Eocene-Oligocene limit. Compared with the only two skulls known of related species (Hyracotheríum vulpiceps and Pachynolophus Iivinierensis), that from Memerlein is distinguished by progressive characters affecting diferent regions but most particularly the braincase; it is not possible, however, to isolate within this evolution the part which leads to a systematic differentiation. Modernization is translated by a considerable increase in size of the braincase, principally in the frontal region, a development of the facial region with anterior displacement of the dental series and a greater specialization of the masticatory apparatus. This evolution parallels the history of the Equidae of the North American early Tertiary, but certain particularities, the form of the alisphenoid, the presence of an anterior frontal foramen, and the structure of the paroccipital apophysis, testifies to the independance of the European forms. 

A lot of isolated teeth of a pteropodid fruit bat has been recently found within an assemblage of micromammals recovered from a karstic fissure filling named Lo Fournas-Il near the locality of Baixas (Pyrénées-Orientales, France). The fauna is Middle Miocene Serravallian age. The fossil fruit bat appears morphologically close to Rousettus; its size is that of a recent medium-sized fruit bat. While the fruit bats are very poorly known as fossils, this discovery shows that one of their recent types of dentitions was perfectly established by Middle Miocene times, and supports the presumed long geologic story of the suborder. One of the major invasions of the Old World fruit bats, supposed originated from SE Asia, reached up to Europe. A suborder unit is added to the miocene fauna of this continent. 

An upper giraffid premolar without any indication about its origin is preserved at the Montpellier University among numerous fossils from the ruscinian formation of Montpellier. It can be related to Samotherium, of the Upper Miocene in Eastern Europe, North Africa and Asia, or more probably to Bramatherium  or Hydaspitherium of the Pliocene of South East Asia. The sedimentological study of the matrix shows a calcareous background, which may indicate that this tooth does not come from the Montpellier formation. 

The first part of this study was devoted to a descriptive analysis of teeth morphologies among the vespertilionine bats. This leads now to a tentative synthesis, providing views on the systematics of the group. The results could be seen according to three distinct but closely related purposes : 1 - the sorting of the genera contents in order to conform the genera units to homogeneous taxa that could represent natural issues of evolutionary lineages ; 2 - the investigation of relationships between extant genera in order to infer the possibilities of common origin ; 3 - according to the preceeding items and to the observed evolutionary trends, a tentative phylogeny, modest and cautious. The contents of many genera are sorted : Leuconoe is removed from subgeneric to generic position, whereas Myotis becomes a subgenus of it ; the myotodont species are cleared away from the Pipistrellus genus ; Glischropus and Scotozous are synonymized within Pipistrellus ; Hypsugo is raised to the generic level ; some species previously ranged within Pipistrellus will form provisionally a collective group, Attalepharca nov. ; the Eptesicus genus is broken up, the excluded species being grouped within Nycterikaupius gen. nov. ; the Nycticeini tribe is defined again after exclusion of Otonycteris , Scotoecus, Scotophilus , and addition of Hesperoptenus ; the species la io and Pipistrellus tasmaniensis are removed to Eptesicus (n.s.) and Pipistrellus dormeri to Scotoecus. Groupings of genera are stated according to the main evolutionary trends of I1/. The relevance of these is often warranted by close morphologic similarities of other teeth. This leads to a recognition of the major evolutionary radiations which occurred in the group. The filiations schematized at the end of the work show the dental relationships observed between the extant genera, and could represent a phylogenic framework. Two major facts are to be underlined : 1- the early divergence of leuconoids ; 2 - the successives crossings to myotodonty from the nyctaloid flow. Fossil data from the literature are punctually and tentatively incorporated within phylogenic sketches. 

Fourteen distinct phyletical lineages which belong at least in three families: Ischyromyidae ALSTON, 1876, Gliridae THOMAS, 1896 and Theridomyidae ALSTON, 1876, have been identified after the study of more than 3600 rodent dental remains from about twenty Early and Middle Eocene european localities. A systematical and phylogenetical revision of these rodents has been achieved. Nearly all the specific and generic diagnosis are emended. Several new combinations and synonymies are proposed. Four new species and two new genera, Euromys nov. (Ailuravinae) and Hartenbergeromys nov. (Microparamyini), are named and described. Euromys nov. gen. is known by three distinctive ypresian (MP 7 to MP 10 european reference levels) chronospecies. This new lineage is thought to be the direct ancestor of Meldimys MICHAUX, 1968 and Ailuravus RUTIMEYER, 1891. A new species of the genus Plesiarctomys BRAVARD, 1850, Pl. lapicidinarum from Condé-en-Brie (MP 8-9 reference level), allows to relate the Plesiarctomys lineage to the Pseudoparamys MICHAUX, 1964 one. The taxa Sparnacomys HARTENBERGER, 1971, Pantrogna HARTENBERGER, 1971, and Corbarimys MARANDAT, 1989 are erected to genus rank; the last one is not thought to be an Ischyromyidae. A new chronospecies of Pantrogna, P. marandati nov. sp. from the locality of Prémontré (MP 10 reference level), is described. This lineage is at the origin of two others, namely Masillamys TOBIEN, 1954, including M. mattaueri (HARTENBERGER, 1975) nov. comb. (MP 10 reference level), and Hartenbergeromys nov. gen., known from MP 10 (H. hautefeuillei nov. sp.) and MP 11 (H. parvus TOBIEN, 1954) reference levels. The phylogenetical position of Hartenbergeromys nov. gen., at the origin of the european family Theridomyidae, is discussed. The systematical and phylogenetical status of two probable Paramyinae, "Paramys" woodi MICHAUX, 1964 and an unnamed genus and species, are discussed. New populations of the primitive Gliridae Eogliravus HARTENBERGER, 1971 and of the primitive Theridomyidae Protadelomys HARTENBERGER, 1968, are described and assigned to previously known species.

  

The composition of assemblages of lizards and Amphisbaenian from the European Eocene are described. At least ten lizard families are identified from the lower European Eocene levels. Eight are still recorded in the last level (Escamps) of the late Eocene. Agamid lizards (genus Tinosaurus) died out by the end of the lower Eocene and Varanid lizards (genus Saniwa) disappeared by the beginning of the late Eocene. Amphisbaenians are recorded throughout the Eocene in Europe. The lacertilian fossil record of Europe and North America show a high degree of faunal resemblance in the early Eocene, followed by a decrease during the later part of the epoch. The lacertilian and amphisbaenian faunas from the European Eocene are not subject to great variations during the period; this is in contrast with the mammal record at the same time. It is argued that the low metabolic rates and the ectothermy of lizards could explain those differences, along with the increasing insularity of the West European area during the late Eocene time.

  

The utilization of screen-washing and attack by dilute acetic acid has permitted the collecting, in the Miocene of the department of Hérault (France), of a very rich ichthyofauna. This fauna is presently comprised of about 60 studied species, of which 11 are new, and represents, in the present state of knowledge, the most varied Miocene selachian fauna described in the world.
The abondance of material has allowed an overall revision to be made; it has thus been possible to complete the description and the figuration of species that were poorly known until now, and to synonymize species that were established on simple morphotypes. Paleo-ecologic study of the ichthyofauna has permitted conclusions to be drawn relative to climate and bathymetry; it was thus possible to show that the Miocene fauna of Hérault was a fauna of a subtropical sea, essentially neritic with rare pelagic contributions.
Knowing the stratigraphic position of the localities, it has been possible to distinguish three faunal assemblages based on associations of species. Some hypotheses on the evolution of certain lineages have been expressed.
The comparison of this fauna with that of other regions permitted the relationships of two diflerent faunal provinces to be specified: the first belongs to the northern domain, characterized by a fauna still subtropical but with numerous temperate water elements; the leoond belongs to the Mesogean domain characterized by warm water forms. It has also lhovm that contemporary faunas could be very different according to the bathymetric zone in which they lived, which furnishes valuable information for the paleogeographic reconstruction of sedimentary basins.
  

A strong lophodonty, an extreme heterodonty, some hypsodonty and regular overlayings of coronal cement are prominent features of the genus Leptolophus (Palaeotheriinae = Palaeotheriidae s.s.). The histological pattern of the teeth unusually joins type II enamel prisms, characteristic of advanced ungulates, together with archaic features, such as an almost complete lack of Hunter-Schreger zonation and a weak expanse of peritubular dentine. The skull is narrow and slender, with an elongated ante-orbital facial region, a moderately notched nasal aperture, a rather elongated post-canine diastem, parallel zygornatic arches and a fairly dorsally located squamoso-mandibular joint.The functional analysis brings to light "ectolophodont" masticatory cycles with two phases, in which maximum power was applied, contrary to equíds, on hindmost teeth; likewise, skull accomodations to increasing height of the teeth are quite different. This study leads to the assumption that Leptolophus may have been light mammals, living in rather open surroundings, browsing on herbaceous plants or leaves cropped close to the ground. Moreover, it appears that it could have been some inadequacy of dental structures to the dietary, which leaded to quick wear of the teeth and to many enamel notches, but had been somewhat balanced by the early increase of hypsodonty, not induced in such a case by a biotop deterioration (as it will happen at the end of the Eocene). This ínadaptation might account for the short duration of the genus Leptolophus, whose the 3 species, L. stehlini, L. nouletí and L. magnus n. sp. are indeed confined in the level MP 16. Its geographical spreading (as far as known, South of western Europe) and the morphological pattern of its dentition suggest that this genus would have been related to early upper Eocene endemic spanish forms.





  

A classification should be both usable and useful,not too complex either in the amount of splitting or in the number of hierarchies involved, and not so simple as to give a false assurance of knowledge of relationships. Classifications are only possible because we do not have complete knowledge of the evolution of the organisms concerned because gaps in the record are necessary to allow the separation of the various taxa. Rodent classification is complicated by the large number of organisms involved and by the geat amount of parallelism that has taken place in the evolution of any and all features. If several independent features are characteristic of a certain taxon, should an effort be made to define the group on the basis of all the features, or should only one be selected as the determinant ? Unless the evolution of the several features was closely linked, the former solution will sooner or later lead to insurmountable problems.