The present study of Artiodactyla from the Middle Eocene of the Geiseltal lignite beds concems systematics, biostratigraphy, and palaeoecology on the basis of 174 specimens: isolated remains to more complete skeletons. Instead of the formerly known five species of two families are now recognized 14 species of the Diacodexeidae, Dichobunidae, Cebochoeridae, and Haplobunodontidae. New species are Aumelasia maniai, Anthracobunodon neumarkensis, Masillabune franzeni. Four species of the Geiseltalfauna are definitely known from elswere, and five species are closely related to those from other European localities. Evidently the faunal situation of artiodactyls during the Middle Eocene of Europe was largely uniform. The distribution of artiodactyls within the sequence of the Geiseltal strata corroborates the biostratigraphical concept of the land mammal age Geiseltalian (Franzen & Haubold l986b) as well as the mammalian reference levels MP 11-13 (Franzen 1987). Reconstructions of the skulls and skeletons allow conclusions on the functional morphology and palaeoecology of the artiodactyls of the European Middle Eocene 

The Palaeobatrachidae until now have not been reported from France. The present note makes known the discovery of very significant remains in three Tertiary localities of France: Cernay (late Paleocene), Laugnac (late Aquitanian) and Sansan (middle Helvetian). The osteologic characters, as well as the geographic and stratigraphic distribution of the family are re-stated with added details. The Palaeobatrachidae are a European family known from the late Jurassic of Catalonia (publication in course by J. Seiffert) to the Plio-Pleistocene (pre-Mindel) of Poland and Rumania. By their way of life and certain osteologic characters they are similar to the Pipidae, but they are distinguished by profound differences. It would be rash to include them in the Pipoidea or even in the Aglossa. They are apparently an independant group, essentially holarctic, which held in Europe a role equivalent to that played by the Pipidae south of Tethys. 

Extensive sampling of the Siwalik deposits of the Potwar Plateau of northem Pakistan and from the Zinda Pir dome of central Pakistan has produced a fossil record of Miocene ctenodactylids that can be correlated with the paleomagnetic time scale. The early Miocene Prosayimys flynni (n. gen., n. sp.) is recognized as the first ctenodactylid in the Indian subcontinent. Prosayimys is ancestral to Sayimys. From the late early Miocene to the early late Miocene, there is an anagenetic succession of three species of Sayimys: S. cf. S. intermedius, S. sivalensis, and S. chinjiensis (n. sp.). Sayimys chinjiensis gave rise to the late late Miocene S. perplexus. A second lineage is represented by Sayimys minor, S. sp. A, and S. sp. B. 

The discovery in the Turolian (Late Miocene) of Dytiko 3 (Macedonia, Greece) of a complete skull with mandibles and cervical vertebrae, atlas and epistropheus, is a very important contribution to the knowledge of the subfarnily Chalicotheríinae. After the description, the comparison with other specimens of Miocene chalicotheres permits the revival of the generic name Macrotherium with a new species M. macedonicum. This genus is mainly characterized by a short snout and an inflated cerebral skull. It coexists during the Miocene with Chalicotherium. A cladistic analysis leads to conclusion that the species which has been described from the Early Middle Miocene of Rusinga must be identified as the type-species of a new genus: Butleria.





  

The study of endocasts, or casts of the endocranial space, have played an important role in shaping understanding of mammalian, and particularly primate, brain evolution. Recently, the reconstructions of three-dimensional virtual endocasts from high-resolution computed tomography images have allowed for the visualization and quantification of endocasts in several Paleocene and Eocene primate species. Here we present the virtual endocast of MaPhQ 289 (informally known as the Montauban 9 skull), a specimen of Necrolemur antiquus Filhol 1873, a middle to late Eocene European primate of the family Microchoeridae. The virtual endocast of MaPhQ 289 reveals a lissencephalic surface morphology with expanded temporal poles and minimal overlap of the cerebellum or olfactory bulb by the cerebrum, which closely resembles the morphology of the endocast of its contemporary relative, Microchoerus erinaceus (Primates, Microchoeridae). MaPhQ 289 yields an endocranial volume (ECV) of 2.36 cm3, about 60% smaller than the volume of the most commonly cited ECV of N. antiquus. Thus, the size of the brain of N. antiquus relative to its body size is likely to be smaller than has been reported in previous literature, highlighting the importance of corroborating older ECV estimates with new evidence using 3-D imaging techniques. 

Bats are among the most numerous and widespread mammals today, but their fossil record is comparatively meagre and their early evolution poorly understood. Here we describe a new fossil bat from dental remains recovered from late Early Eocene sediments at Prémontré, northern France. This 50 million-year-old bat exhibits a mosaic of plesiomorphic and apomorphic dental features, including the presence of three lower premolars, a single-rooted p3, short p4 with metaconid, myotodont lower molars and a tall coronoid process of the dentary. This combination of features suggests it is an early member of Vespertilionidae, today’s most speciose and geographically widespread bat family. The Prémontré bat has bearing on hypotheses about the origins of vesper or evening bats (Family Vespertilionidae), as well as crown-group chiropterans.


  

A new hypothesis for the phylogenetic relationships of recent anomalurids and graphiurids is proposed, based on information from evolutionary lineages of Paleogene European rodents, particularly Gliridae, and Eocene Algerian Zegdoumyidae. Differences in first occurrences, in paleogeography, and in infraorbital structure in glirids (protrogomorphy and pseudomyomorphy) and graphiurids (hystricomorphy) separate Graphiuridae from Gliridae (Graphiurinae is here raised to family rank). Similar considerations, and dental morphology, suggest that Anomaluridae (appearing in the late Eocene) and Graphiuridae (appearing in the Pliocene) are related to early Eocene Zegdoumyidae. 

The study of the bovids from Al Jadidah (Hofuf Formation, Saudi Arabia) confirms that the fauna comes from a pre-Hipparion level. The Al Jadidah age is close to that of Fort Ternan (14 m.y.) and Beni Mellal, but cannot be older than that of Fort Ternan. The age of the Hofuf Formation is close to but slightly older than the oldest deposits of the Ngorora Formation (Kenya). 7 to 9 species have been recorded, of which 2 to 4 remain indeterminate. If the great specific diversity of te bovids from this locality gives evidence of immigrations from anterior Asia (Turkey) (e.g. Pachytragus Iigabuei sp. nov.), the bovid assemblage of Al Jadidah results in fact from a double influence: from the anterior Asia and mainly from Africa (e.g. the Caprotragoides lineage and the Neotragini? Homoiodorcas). The Al Jadidah bovids reflect, on the whole, the predominant character of open to very open environment, which supports the conclusions drawn from our two preliminary studies. 

Is hereby described an impressive lchnoiauna belonging to the Lower to Middle Ludian of the Gard (S. France). The slab, already cleaned over a length of 18 m, is located near the top of the Potamides aporoschema lacustrine limestone (Lower Ludian, Euzet zone). It is therefore older than the Célas sandstone deposit, and still more than the Melanoides albigensis and M. acutus marly limestone corresponding to the Upper Levels of the Ludian stage. Although biostratigraphically older than the La Débruge and Montmartre zone, the biotope shows already a sampling of very tall Artiodactyles, Perissodactyles and Carnivorous. One of the most « majestic ›› Artiodactyles, Anopolotheriipus lavocati, nov., points out a huge size type. To mention also among the Ichnotypes described, 10, the big Perissodactyle Palaeotheriipus similimedius, nov., and the big Carnivorous Hyaenodontipus praedator, nov. 

The review of material recently collected in the new localities from the “Phosphorites du Quercy", and different localities from the South of France, bring new informations on the genus Eucricetodon THALER. 1966, and Pseudocricetodon  THALER,  I969 (Middle and Upper Oligocene. Western Europe). Thanks to Eucricetodon huerzeleri VIANEY-LIAUD, 1972, which were unsufficiently known until now, is proposed. During the middle Oligocene Eucricetodon atavus  MISONNE, 1957 seems to give rise to two lineages. One of them led to Eucricetodon huberi,which however exhibits a larger size and a development of progressive characters on the teeth. The other would be Eucricetodon huerzeleri well differentiated at the “Mas de Pauffié" standard level (beginning of the upper Oligocene). The ornementation of lower incisors is described, when possible. Though the fossils are not abundant, it seems that the ancestral lineage, Eucriretodon atavus, remains in  the upper Oligocene (Boningen standard level). evolving into Eucricetodon praecursor SCHAUB, 1925 (Rickenbach standard level). The characters of Eucricetodon dubius  (SCHAUB. 1925), represented by a numerous population in Pech Desse and Pech du Fraysse (Quercy). confirm that this species and Eucricetodon praecursor  belong to two different lineages. As Eucricetodon dubius shows more primitive features, this species could not originale from Eucricetodon atavus -Eucricetodon huberi. The appearance of this species at the level of Mas de Pauffié could be the result of an immigration. A new definition of Pseudocricetodon incertus (SCHLOSSER. 1884) is given. This species has been found in several localities, where it had not been identified until now. lts comparison with Pseudocricetodon moguntiacus  (BAHLO. l975), found at several localities from the standard level of Antoingt (end of middle Oligocene). shows a parallel evolution to that of Pseudocetodon incertus, which is  of larger size and with a less complicated pattern of teeth. 

Some fragmentary remains of spines and endoskeletal elements, referred here to as ctenacanthid like elasmobranchs, are recorded in the Middle Devonian of Bolivia. These specimens, and some others from the Eodevonian of Brasil represent the only  Devonian fish remains hitherto known from South America and indicate that further discoveries may be made in their original localities respectively.
  

Near "la Lieude", in the Lodève basin, more than a thousand of footprints are distributed in a twenty of trackways which amounts to 220 m length. They have been found on calcareous siltstone level in the B site named also "Réserve Naturelle Volontaire". This last is located in the Saxonian summit dated Upper Permian. "La Lieude" tracks are described by using statistical methods then they are compared with others from the world Permian. What allows to distinguish 4 following ichnotaxa: Lunaepes ollierorum nov.ichnosp., Merifontichnus thalerius nov. ichnogen. and nov. ichnosp., Planipes brachydactylus nov.ichnosp. and Brontopus circagiganteus nov. ichnosp. All these traces are attributed with possibility or probability to Therapsida or to Therosauria, except Brontopus circagiganteus nov. ichnosp. that could be due to Caseamorpha. All these animals whose sizes have been estimated between l and 5 m lived probably in a playa environment.The biological and sedimentological data from "la Lieude" footprints levels compared with informations provided by the tracks orientations, suggest the following scenario. Animals coming from the North have crossed a sandy channel bank with plants zones by directing to the South for the majority. Maybe, they were going to the lacustrine part of the playa, close to "la Lieude" footprints that they have just trampled on. 

The first part of this study was devoted to a descriptive analysis of teeth morphologies among the vespertilionine bats. This leads now to a tentative synthesis, providing views on the systematics of the group. The results could be seen according to three distinct but closely related purposes : 1 - the sorting of the genera contents in order to conform the genera units to homogeneous taxa that could represent natural issues of evolutionary lineages ; 2 - the investigation of relationships between extant genera in order to infer the possibilities of common origin ; 3 - according to the preceeding items and to the observed evolutionary trends, a tentative phylogeny, modest and cautious. The contents of many genera are sorted : Leuconoe is removed from subgeneric to generic position, whereas Myotis becomes a subgenus of it ; the myotodont species are cleared away from the Pipistrellus genus ; Glischropus and Scotozous are synonymized within Pipistrellus ; Hypsugo is raised to the generic level ; some species previously ranged within Pipistrellus will form provisionally a collective group, Attalepharca nov. ; the Eptesicus genus is broken up, the excluded species being grouped within Nycterikaupius gen. nov. ; the Nycticeini tribe is defined again after exclusion of Otonycteris , Scotoecus, Scotophilus , and addition of Hesperoptenus ; the species la io and Pipistrellus tasmaniensis are removed to Eptesicus (n.s.) and Pipistrellus dormeri to Scotoecus. Groupings of genera are stated according to the main evolutionary trends of I1/. The relevance of these is often warranted by close morphologic similarities of other teeth. This leads to a recognition of the major evolutionary radiations which occurred in the group. The filiations schematized at the end of the work show the dental relationships observed between the extant genera, and could represent a phylogenic framework. Two major facts are to be underlined : 1- the early divergence of leuconoids ; 2 - the successives crossings to myotodonty from the nyctaloid flow. Fossil data from the literature are punctually and tentatively incorporated within phylogenic sketches. 

A recent campaign of excavations (1965-68) undertaken by the Laboratoire de Paléontologie of Montpellier in pockets of the Quercy phosphorites, has permitted the dating of several localities thanks to the analysis of their micromammalian fauna.
The rodents of localities belonging to the middle Oligocene (La Sauvetat zone) are the object of this study. This rodent fauna has been compared to the forms coming from three stratified localities belonging to the biochronologic zone of La Sauvetat :  Balm, Montalban and Lovagny.
The abundance of material, notably of theridomyids and cricetids, permitted a summary statistical study to be effected and gave some precise details on the variability of the species (Theridomys varians in particular). Other details were provided on certain groups, notably on the upper teeth of Scíuromys cayluxi and on the milk teeth of theridomyids. Some specimens of «Sciurus» sp. and of Plesispermophilus angustidens were collected. By this means, it has been possible to establish, for the first time, a precise upper limit to the epoch of their appearance in Europe. 

Seventeen species of cricetid rodent are recognized and described from lower and middle Siwalik deposits in the Potwar Plateau of Pakistan. These species are grouped in three categories, characterized as subfamilies (e. g., Megacricetodontinae, Myocricetodontinae, and Dendromurinae); an additional and more abundant category of rodents from these deposits, the Democricetodontinae, is excluded from this study, and will be described in a later study. Fifteen of the species are new, and four new genera are described. The Siwalik cricetid taxa are : Megacricetodon aquilari, n. sp.; Megacricetodon sivalensis, n. sp.; Megacricetodon daamsi, n. sp.; Megacricetodon mythikos, n. sp.; Punjabemys downsi, n. gen. & n. sp.; Punjabemys leptos, n. gen. & n. sp.; Punjabemys mikros, n. gen. & n. sp.; Myocricetodon sivalensis, n. sp.;  Myocricetodon sp.; Dakkamyoides lavocati, n. gen. & n. sp.; Dakkamyoides perplexus, n. gen. & n. sp.; Dakkamys asiaticus, n. sp.; Dakkamys barryi, n. sp.; Dakkamys sp.; Paradakkamys chinjiensis, n. gen. & n. sp.; Potwarmus primitivus, n. gen.; and Potwarmus minimus, n. gen. & n. sp. This diverse record of middle Miocene small mammals illuminates a profound radiation of cricetid rodents in southem Asia, the effects of which were felt in Europe and Africa as well as the rest of Asia. 

Based on material from 30 localities, morphologic dental, cranial and biometric analyses have permitted the characterization of two parallel Issiodoromyine lineages, and also the definition of diverse species representing several evolutive stages. Thus it is that new lineages complete the contribution made by the Theridomyinae and Cricetidae and permit, for the Quercy in particular, additional precision in the biochronologic succession of the localities. One of the lineages is limited to the genus Pseudoltinomys LAVOCAT; the other evolves from the genus Elfomys HARTENBERGER to the genus Issiodoromys BRAVARD in GERVAIS. The latter is affected by profound anatomical changes due to a functional modification of the mastication apparatus. These changes seem to be able to be put in relation with the aridification and cooling of the climate at the end of the Eocene. At the end of the middle Oligocene, a new chewing structure is achieved. It is found in diverse living rodents that inhabit a rather arid steppe environment (Cavia, Pedetes, Ctenodactylus). To these supposed nearby ecologic conditions, these rodents have responded in a convergent fashion. It is possible to attribute to the extreme specialization of Issiodoromys its incapacity to adapt to the new climatic crisis of the end of the Oligocene. The arrival of immigrants may be considered as another cause of its disappearance at this time, complementary or not with the first. 

The object of this work is the study of the artiodactyls of the karstic locality of Lissieu near Lyon. Among the sixteen species counted, five (Hyperdichobune nobilis, Hyperdichobune sp., Haplabunodon mulleri, Tapirulus depereti and Pseudamphimeryx schlosseri) are cited for the first time in this fauna. The description of part of the anterior dentition of «Dichobune» langi permits the referal of this species to the genus Hyperdichobune. The new selenodont species Dichodon lugdunensis  described in this work belongs, with its ancestor Dichodon simplex, to a lineage distinct from that of Dichodan cartieri and Dichodon ruetimeyeri. Finally, the taxonomic reality of the Lissieu species, Lophiobunodon rhodanicum Deperet, is justified. All the other species have been compared with those of Egerkingen. Even though with most of them no difference in age can be shown between the two deposits, some however indicate clearly by their degree of evolution that Lissieu is slightly more recent than Egerkingen. ln the history of Eocene mammals, the Lissieu fauna, because of its homogeneity, can be considered as a sure reference fauna of the Auversian period. 

The fossil mammals discovered in the quarry of Rians (Sparnacian, Provence) are described. Among these forms, Hyracotherium is interesting because of the little molarization of the lower premolars and its small size, and Diacodexis by its small size and very primitive astragalus ; they may be the most primitive representatives of their respective orders. Also, Proviverra eisenmanni n. sp. is the smallest and most primitive hyaenodontid yet described. Hyopsodus itinerans is the first species of this genus described France. Among other rare fossils is a new species of bat, a small palaeoryctid, and other forms not yet identified. Marsupials are varied. Several new species are present among the rodents. The fauna is well-balanced and rich in small hyopsodontid condylarths. It is stratigraphically situated at the
Dormaal reference-level, at the base of the early Eocene, and is considered equivalent to the late Clarkforkian of North America. The hypothesis is presented that new forms appearing at the beginning of the Wasatchian in North America migrated, in fact, at that time from Europe.

  

A new European rodent from the middle Eocene of Spain, Zamoramys extraneus n. gen., n. sp., appears to be closely related to the middle Eocene chapattimyid rodents of Indo-Pakistan. This contradicts the generally accepted paleobiogeographic hypothesis of a Tethyian barrier between Europe and Asia isolating Europe during the middle Eocene. Because of this barrier, some authors have proposed that European and Asian rodents were not closely related, their similarity being the result of morphological convergence. Here monophyly has been tested, using the parsimony criterion, based on an analysis of dental characters (including discussing of homology and the validity of some characteristics). Our results indicate a phylogenetic relationship among the Asiatic Ctenodactyloidea, Zamoramys from Spain, and the European endemic Theridomyoidea. We also conclude from our analysis that theridomyoids and European ischyromyoids are probably not closely related phylogenetically. 

A fossil species of the extant genus Leuconoe, L. lavocati n. sp. from Le Garouillas Oligocene locality, Quercy phosphorites, SW-France, is established in nomenclatural standards.