The rodent fauna of Beni-Mellal is characterized by the abundance of ctenodactylids and cricetids. The latter are represented by four distinct species, among which a new form. Dakkamys zaiani nov. gen., nov. sp. is described. A detailed morphological analysis shows that, contrary to that which had been established before, « Cricetodon ›› atlasi Lavocat, 1961, is not closely related to any European form known; this species is attributed, in consequence, to the new genus Mellalomys. A simple biometric analysis has shown that the genus Myocricetodon Lavocat, 1952, is represented in this locality by two distinct species. The systematic homogeneity of the Beni-Mellal cricetids is also demonstrated: they can, as a matter of fact, all be referred to the subfamily Myocricetodontinae. The definition of this subfamily is completed. The sciurids and glirids are also reviewed in the light of new systematic and biogeographic information established ln Europe. A new species of Atlantoxevus from the early Pleistocene of Morocco, A. huvelini nov. sp., is described. It is probably the descendant of A. tadlae from Beni-Mellal. Biogeographic analysis leads one to consider this fauna as the result of geographic isolation in the Maghreb since the late Oligocene or the early Miocene. In particular no direct European influence can be discerned. Stratigraphic considerations resulting from the discovery of new localities in North Africa lead to the confirmation of the ante-Vallesian age of this fauna and to its parallelism with the faunas of La Grive in Western Europe and Fort Ternan in East Africa. The peculiar geologic nature of this locality is discussed.
  

Le matériel provenant de nouvelles fouilles dans les phosphorites du Quercy, soumis aux divers spécialistes, a conduit à la soutenance de plusieurs thèses d'état: sur les rongeurs (Hartenberger, Vianey-Liaud), les Chiroptères et Insectivores (Sigé), les Artiodactyles (Sudre), les Squamates (Rage) et, en partie, les Chéloniens (De Broin). Chacun dans son domaine, à côté des conclusions d'ordre évolutif, paléogéographique ou paléoécologique, a pu établir des successions stratigraphiques des gisements du Quercy qui se sont révélées largement concordantes. Ainsi la succession des faunes du Quercy est actuellement l'une des mieux établies. Dans ce contexte, les différents spécialistes ont décidé de conjuguer leurs efforts dans l'étude monographique de plusieurs gisements repérés le long de cette échelle, afin de rassembler l'information paléontologique sur des faunes bien précises et de tenter d'obtenir des indications d'ordre taphonomique. 

The description of original material of anthracothere and proboscidean in the new locality of Bir el Ater 3 from East Algeria, and a thorough review of early Libycosaurus remains of Bir el Ater 2 allows us validating L. algeriensis as the smallest and earliest species of Libycosaurus and probably the earliest migrant of the genus from Asia. The presence of a Tetralophodon in the Neogene Nementcha formation might represent the earliest occurrence of the genus in Africa. These original fossil remains allow us to discuss the age of the Neogene part of the Nementcha formation close to the Serravalian/Tortonian boundary. 

Le Quercy est aujourd'hui un vaste plateau calcaire, parcouru par un réseau karstique actif, pro· fondément entaillé par des vallées aux falaises abruptes, comme celles du Lot ou du Célé. Sur un sol peu épais domine la forêt de chênes, accompagnés de cornouillers, érables, genévriers. La faune est pauvre, peu diverse, et les nombreux chasseurs se satisfont de gibier d'élevage ...
Il y a trente-cinq millions d'années environ, le paysage était bien différent. La période de l'Eocène supérieur, qui s'achevait, avait été chaude et humide, si l'on se réfère à la fois aux paléotempératures (calculées à partir de sédiments marins extra-européens) et aux restes fossilisés de végétaux typiquement tropicaux.
Le Causse du Quercy devait être un plateau très disséqué par la karstification, à surface lapiazée creusée de gouffres en rapide évolution interne, et couvert d'un sol assez épais. Une forêt tropicale humide, avec notamment des Myricacées et des gymnospermes, recouvrait l'ensemble du pays, à peine interrompue au niveau des rares points d'eaux situés dans les bas-fonds. Dans ce cadre vivait une faune considérablement différente de la maigre faune actuelle. Si nous pouvons l'imaginer, c'est grâce aux cadavres des animaux entraînés dans les cavités par les eaux courantes ou les prédateurs, ou bien logeant et mourant dans les milieux souterrains. Nombre de ces restes, fossilisés, sont parvenus jusqu'à nous et sont aujourd'hui l'objet d'étude. 

The status of the giant bird taxa Liornis floweri and Callornis giganteus from the Santa Cruz Formation (late Early Miocene) of Patagonia, first described by Ameghino (1895) is reassessed on the basis of a re-examination of the type material at the Natural History Museum, London. Liornis floweri, which lacks a Pons supratendineus on the tibiotarsus and has an unbifurcated Canalis interosseus distalis on the tarsometatarsus, is clearly a brontornithid and is considered as a junior synonym of Brontornis burmeisteri. Ameghino’s replacement of Callornis by Eucallornis is unjustified. Callornis giganteus is a chimera based on a phorusrhacid tarsometatarsus (probably belonging to Phorusrhacos longissimus) and a brontornithid tibiotarsus. The latter can be considered as the lectotype of Callornis giganteus, which may represent a small morph of Brontornis burmeisteri or a distinct taxon. It is referred to here as Brontornithidae indet. The tarsometatarsus described by Dolgopol de Saez (1927a,b) as Liornis minor and considered by her as a gracile brontornithid apparently has a bifurcated Canalis interosseus distalis and should therefore be placed among the Phorusrhacidae. 

The Las Águilas site near Porvenir de Jalpa, Coahuila, Mexico, is extremely rich in tetrapod remains comprising both bones and trackways of several dinosaur taxa of late Campanian age. Within a 50 m thick section we identified at least nine layers with dinosaur bone assemblages. In one of these the dinosaur bones are associated with remnants of eusuchian crocodilians, turtles, plesiosaurs, pterosaurs, tyrannosaurids, dromaeosaurids, parksosaurid, hadrosaurids, ceratopsids, and ankylosaurs. This layer is also rich in coprolites of turtles, crocodilians and likely theropods, thus providing evidence for the wealth of Late Cretaceous vertebrate life in the area. 

Lalieudorhynchus gandi gen. nov. and sp. nov. is a new caseid synapsid from the Permian of the Lodève Basin, Occitanie, France. This new taxon is represented by a partial but well-preserved postcranial skeleton, and is characterized by the following apomorphies: a transverse section of the sacral and anterior caudal neural spines with a very thin keel-like process anteriorly, a slender dorsal tip of the dorsal and caudal spines, a narrow distal end of the first sacral rib, a fossa on triceps process of metacoracoid, and a very large distal tarsal 1 of same width than the astragalus, with nearly all sides being shallowly concave.
The skeleton corresponds to a sub-adult individual that was excavated from the La Lieude Formation dated as Roadian-Capitanian (Guadalupian). A sedimentological and taphonomical analysis of the type locality, together with preliminary osteohistological observations, suggest that this new French caseid was rather aquatic, as already hypothesised for other large forms.
A phylogenetic analysis of caseids is performed to test the position of this new taxon and to better understand the evolution of the clade: interestingly, Lalieudorhynchus gandi gen. nov. et sp. nov. is closer to the NorthAmerican “Cotylorhynchus” hancocki than to the other French caseids Ruthenosaurus and Euromycter from the Artinskian of the geographically closer Rodez Basin. These two last caseids document the Artinskian radiation of the clade, which remained diverse until Olson’s extinction. Caseids survived, as Lalieudorhynchus is one of the youngest representatives of the clade, and may have used novel ecological strategies to access their vegetarian food sources.

  

This paper presents the description and the interpretation of recently discovered traces on a Lower Hettangian dolomitic outcrop in the Bédarieux area, Southern France. One trace set immediately attracted the attention by its resemblance to a small sauropod pes-manus couple but no trackway was visible. As the other traces have a variety of shapes with no obvious significance, it took a thorough examination of the 3D and sedimentological data to come to the conclusion that most traces likely were sauropod tracks made under diverse conditions. Sedimentological and ichnological data indicate that the tracks have been made in the intertidal zone of a carbonated tidal flat shortly before an emersion period. It appears that that the variety of trace shapes is due to a variety of water depths: the sauropods were punting when the water level was high. The lack of trackways seems due to the combination of an underprint situation, buoyancy effects and the small size of the track-bearing slab. Several hypotheses can be considered for explaining the very small size of the tracks, such as insular dwarfism or the immaturity of the trackmakers. 

Macrauchenia patachonica was among the largest litopterns. It had a long neck with elongated cervical vertebrae, unique among endemic South American ungulates. We calculated the pattern of stress in the joints between the vertebral centra along the neck of the recently-extinct litoptern mammal M. patachonica for various hypothetical neck postures to determine which one is optimal. We also determined the zygapophyseal alignment positions for the neck, assuming a wide range of values for the thickness of the intervertebral discs. We concluded that a vertical posture is the one that best meets the requirements of nearly constant stress. This upright posture was probably a frequently adopted posture by M. patachonica while feeding or standing. It is also possible that occasionally it could adopt a gerenuk-like posture. In almost any other position, the standard deviations of stress values (SD) divided by mean stress (MS) have values between 0.4 and 0.5. Since it was a mixed feeder, M. patachonica probably used different postures to reach resources at different heights. However, an almost horizontal posture was required for the optimal articulation of the neck vertebrae. It probably represents the posture during fast locomotion, as suggested in a previous biomechanical study of locomotion. 

Sirenians are among the common marine fossil remains found in the Faluns deposits of western France. We describe new material of a Dugongidae sirenian from the middle Miocene Savignean facies of La Morfassière quarry (Indre-et-Loire, northwestern France) that includes a well preserved and almost complete skull associated with its mandible, several vertebrae and ribs. The cranial remains exhibit features that allow to attribute them to Metaxytherium medium, a species recorded from the middle and early late Miocene of European and Mediterranean coasts. The discovery of an associated skull and mandible of this species is unusual in this area and deserves to be reported, mostly because its preservation contributes to a better knowledge of the variable structure of its skull and teeth anatomy. For the first time the body size and weight of M. medium can be assessed through regression equations based on skull measurements. The particularly good condition of the material can be explained by the calm and deep marine environment in which it was deposited.
  

Aside from Madtsoiidae, anilioids, and Boidae that were studied previously, the middle Palaeocene of ltaborai (BraziI) has produced Ungaliophiinae ("tropidophiids"), booid-grade snakes incertae sedis, and a possible Russellophiidae (Caenophidia) that are described in the present article. This article is the third and final report on the snakes from the locality. The Ungaliophiinae (Paraungaliophis pricei gen. et sp. nov.) are rare whereas the booid-grade snakes incertae sedis (ltaboraiophis depressus gen. et sp. nov., Paulacoutophis perplexus gen. et sp. nov.) are more frequent. A single vertebra is referred to the Russellophiidae (Caenophidia) with reservation. An update of the whole fauna of snakes from ltaborai is provided. Hechtophis austrinus that was tentatively referred to the erycine Boidae is now regarded as a Boidae incertae sedis. Most snakes from Itaborai are known only from the locality. Astonishingly, only the ailioids Coniophis cf. C. precedens gives possible evidence of interchanges between South and North America. The fauna of snakes from Itaborai, as well as the other Palaeocene faunas of snakes from South America are distinct from those of the Cretaceous and the Eocene of South America; they appear to be more different from the Cretaceous faunas than from those of the Eocene. The fauna from Itaborai is the richest and most diverse assemblage of snakes from the Palaeocene worldwide; it shares only a few taxa with other Palaeocene localities. 

Morphological and biometrical description of skulls, teeth, and limb bones of extant and fossil Old World herniones (including E. hydruntinus) and of New World 'stilt-Iegged' and other slender species from Blancan to Holocene. An Appendix presents ways in which the approximate size of some missing bones or dimensions may be deduced from available ones.

The discussed and/or illustrated fossils were found in Bolivia (Tarija), Canada (Yukon), China (Choukoutien, Gulongshan, Jiling, Loufangzi), Ecuador (Oil Fields), Ethiopia (Melka Kunturé), France (Lunel-Viel), Germany (Süssenborn), Greece (Agios Georgios, Petralona), Hungary (Dorog), Italy (Romanelli), Mexico (Cedazo, San Josecito), Mongolia (Sjara-osso-gol), Spain (Venta Micena), ex-Soviet Union (Akhalkalaki, Binagady, Chokurcha, Chukochya, Kabazi, Kolyma, Krestovka, Kurtak, Staroselie, Tologoj), USA (Alaska, Arkalon, Cedar Meadow, Channing, Conkling, Dry Mountains, Hay Springs, Leisey Shell Pit A, Lissie Formation, Natural Trap, Pool Branch, Powers Ranch, Rock Creek, San Diego, Santo Domingo, Seymour Formation, Shelter, Slaton, Trinity River). Numerous raw or statistically elaborated data are given in Tables.

There is no evidence for the existence of Old World hemiones in the New World nor of 'stilt-Iegged' equids in the Old World. The first 'stilt-Iegged' equid was found at Santo Domingo, New Mexico, and is believed to be Late Blancan. It was probably at the origin of E. calobatus (Arkalon, Rock Creek) and of the smaller E. semiplicatus (Channing, Rock Creek). Slender, but not 'stilt-Iegged', equids found at Natural Trap, Wyoming, ca. 12 ky ago belong to Amerhippus. AlI these species share with Oid World Sussemiones (and some hemiones) peculiar patterns on the lower cheek teeth.

The slender Equus sp. B of Leisey Pit A, Florida, ca. 1.2 Ma, as weIl as Amerhippus francisci and E. tau (probably a senior synonym of E. quinni) share conventional lower cheek teeth patterns. The skulls of A. francisci and E. tau, however, are quite different.

Paleontological data suggest a common origin of Amerhippus, Sussemiones, and 'stilt-Iegged' equids during the late Blancan. Old World hemiones seem to have differentiated later. 

In 2002, Antunes & Cappetta published several new taxa (genera and species) in a paper on Cretaceous elasmobranch faunas from Angola. One of the new genera was named Angolaia (Rajiformes, Hypsobatidae; type-species: Angolaia benguelaensis Antunes & Cappetta, 2002, Late Campanian/Early Maastrichtian of Angola). Recently, Dr. Christian Kammerer kindly informed us that the genus Angolaia was preoccupied by a cicadellid homoptere (Insecta), published by Linnavuori & Al-Ne'amy, 1983. So, according to mIes of the International Code of Zoological Nomenclature (ICZN 1999, articles 52, 60) this name became unavailable. Consequently, the Rajiformes genus Angolaia is a junior homonym, invalid and must be rejected. In replacement, we propose the new name Angolabatis. 

A new carcharhinid shark species, Physogaleus hemmooriensis sp. nov., is described from the Lower Hemmoorian (Behrendorfian, late Burdigalian, early Miocene) of Werder, Lower Saxony, Germany. P. hemmooriensis also occurs in the Edegem and Antwerpen Sands Members of the Berchem Formation, Belgium, and in the Miste Bed, Aalten Member of the Breda Formation, The Netherlands, which have an early to middle Miocene age. In the Western Atlantic region, the taxon is present in the early Miocene Calvert Formation of Delaware, U.S.A, which is largely contemporaneous with the Hemmoorian. 

Eosimiid and amphipithecid primates document a long and significant history of primate evolution throughout the Eocene in Southeast Asia. Despite the absence of a comprehensive post-Eocene fossil record, it was generally hypothesized that both families left no descendant in Asia. Recently, two new small-bodied taxa, Bugtipithecus and Phileosimias, have been recovered in early Oligocene coastal deposits from the Bugti Hills (Balochistan, central Pakistan) and referred to the families Amphipithecidae and Eosimiidae, respectively, on the basis of dental fossil remains. In this paper, we provide more exhaustive description, comparison, and discussion of these taxa. As for tarsiid and sivaladapid primates, the persistence of eosimiids and amphipithecids into the Oligocene clearly demonstrates that low latitudes of South Asia provided a continuous access to tropical refugia during the climatic deterioration characterizing the late Eocene-early Oligocene interval, which was seemingly lethal for primate communities elsewhere across the Holarctic continents. As a contribution to the ongoing phylogenetic debates regarding the position of eosimiids and amphipithecids on the primate family tree, we have performed a cladistic analysis in a high-level primate systematic context in order to assess the position and the role of these new taxa in that phylogenetic issue. Our results support the view according to which eosimiids and amphipithecids (and by extension Phileosimias and Bugtipithecus, respectively) are stem anthropoids. These fossils from Pakistan document an unsuspected Oligocene phase of the evolutionary history of anthropoid primates in southern Asia, which clearly enhances the extent of the anthropoid radiation in this province during the Paleogene. Several phylogenetic and paleobiogeographic aspects are discussed, notably the intra- and inter-relationships between Paleogene Asian and Afro-Arabian anthropoids, and the resulting potential dispersal models between both land-masses during the Paleogene. 

The scientific career of D.E. Russell began with a Pliocene fauna from Oregon, and then turned in the direction of European Paleogene mammals. Field work followed by study of the mammals that were collected, firstly in the Paleocene and later in the early Eocene, greatly rejuvenated learning in this field. Syntheses on the Northwest European Tertiary basin and on European marnmals and stratigraphy came next. Research on the Eocene of Asia was carried out jointly with Gingerich on Pakistan and with Dashzeveg on the faunas of Mongolia. An important synthesis on the entire Paleogene of Asia, joint with Zhai, followed. Field work in Africa with Sigogneau-Russell led to the discovery of Mesozoic mammals there. A synthesis of mammalian paleofaunas of the world was written with Savage, and a similar synthesis of Cenozoic vertebrate faunas is currently being prepared. These achievements reflect the perennial importance of field work, numerous collaborations with both amateurs and professionals, and the human qualities of this author.
  

The rodents from the spanish locality of Can Ponsic 1 bring new data about some rodents species of the beginning of the Upper Miocene in South-West Europe. The criticims made by Mein and Freudenthal about the validity of the species Hispanomys thaleri from Can Llobateres are not justiíied. The study of the anatomy of the skull of Rotundomys from Can Ponsic 1 gives accurate information about the affinity of this genus with Cricetulus, and shows that the hypothesis, according to which Rotundomys is an ancestral form of the Arvicolids, is unlikely. The systematics of Heteroxerus and the phylogeny of the mio-pliocene Muscardinus species are also discussed. The Can Ponsic 1 locality is a little older than Can Llobateres.

  

The rodent-fauna (Cricetidae and Gliridae) recently found at Castell de Barbera (Spain) is similar to those from the other locslities of the Valles - Penedes - Can Ponsic 1 and Can Llobateres - : same composition and similar evolutionary level of the different species. On the other hand this fauna is different from those of Upper Vindobonian and Vallesian localities of the Calatayud - Teruel area. Castell de Barbera has an intermediate chronological position between the localities of Anwil (Switzerland) and Can Ponsic 1. It is still not possible to validate or invalidate the initial attribution based on absence of Hipparion of Castell de Barbera to the Upper Vindobonian. 

A new Devonian vertebrste locality has been discovered in 1976 in the basal part of the Khush-Yeilagh Formation in the eastern Alborz Mountains of Iran. The fossils occur in a band one centimeter thick which is identifiable at other outcrops in the area. A preliminary study of the remains has yielded the following faunal list : Placodermata (Phlyctaeniida indet.. Groenlandaspididae indet.. Coccosteidae indet., Holonema sp., Ptyctodontida indet., Antiarcha indet., Bothriolepis cf. kwangtungensis), Elasmobranchii (Ctenacanthidae indet.), Acanthodii (Gyracanthus sp., « Onchus » overathensis, Ischnacanthiforme indet.), Dipnoi (? Dipteridae indet.), « Crossopterygii ›› (Onychodus cf. sigmoides, Holoptychiidae indet., Osteolepididae indet.). This fauna is older than the Middle Givetian and probably of Middle or Lower Eifelian age. Owing to its diversity, it may prove possible to use it in the future as a reference fauna for the study of the Devonian vertebrates in Central Asia and the Middle East. 

A brief historical account of fossil vertebrate discoveries in the Eastern part of the Paris Basin between the beginning of the nineteenth century and 1950 is given. Other localities discovered since then are presented. A reconstruction of past landscapes is briefly elaborated. A biozonation based on mammals is proposed, from the Late Thanetian to the Middle Bartonian. Paleogeographical considerations are added. Suggestions regarding the search for new marnmal localities are made.