Un crane de Chalicothere (Mammalia, Perissodactyla) du Miocène supérieur de Macédoine (Grèce) : remarque sur la phylogénie des Chalicotheiinae

Louis de Bonis , Geneviève Bouvrain, George D. Koufos and Pascal Tassy

The discovery in the Turolian (Late Miocene) of Dytiko 3 (Macedonia, Greece) of a complete skull with mandibles and cervical vertebrae, atlas and epistropheus, is a very important contribution to the knowledge of the subfarnily Chalicotheríinae. After the description, the comparison with other specimens of Miocene chalicotheres permits the revival of the generic name Macrotherium with a new species M. macedonicum. This genus is mainly characterized by a short snout and an inflated cerebral skull. It coexists during the Miocene with Chalicotherium. A cladistic analysis leads to conclusion that the species which has been described from the Early Middle Miocene of Rusinga must be identified as the type-species of a new genus: Butleria.