The entelodonts of the Quercy phosphorites are essentially known from isolated teeth. The skull remains attributed to Entelodon magnum by Kowalewsky in 1876 and the type left mandible of Enteladon depereti RÉPELIN, 1918 do not belong to this family. From a morphologic point of view, the teeth from Quercy are quite comparable to those from the type localities of E. magnum and E. deguilhemi. In the absence of distinctive anatomical criteria between E. magnum (small species from the level of Ronzon) and E. deguilhemi (large species from the level of Villebramar) they are referred, according to their size, to one or the other of these two species. However, some of them are intermediate in size. Their existence speaks in favor of a close relationship between E. magnum and E. deguilhemi and confers a real stratigraphic interest to these entelodonts of Western Europe. Finally, some anatomical characters (notch on the mesial border of P4, presence of a paraconid joined to the metaconid, trigonid higher than talonid) considered until now as peculiar to the North American Archaeotherium and not yet cited in its European contemporary Entelodon have been brought to light on the teeth from Quercy.

  

The two species of Sandgrouse from Quercy, Pterocles validus MILNE-EDWARDS and P. larvatus MILNE-EDWARDS, are ascribed to the genus Archaeoganga MOURER-CI-IAUVIRÉ which includes a third, very large species, A. pinguis. The sandgrouse of Saint-Gérand-le-Puy, Pterocles sepultus MILNE-EDWARDS, is ascribed to a new genus, Lepzoganga. This form appears in the Upper Oligocene of Quercy, in Pech Desse and Pech du Fraysse localities, and is still present in the Lower Miocene of Saint-Gérand-le-Puy and Paulhiac. Recent sandgrouse live in semidesert or desert areas. The indications provided by mammal and bird faunas in the localities where sandgrouse were found, confirm that the paleoenvironment was open and arid. The morphological study of these fossils indicates that, in the Upper Eocene, the Pteroclidae were already completely individualized with respect to Charadriiformes.
  

The teeth and the astragale of the Creodonta from Ste-Néboule (Lot) are referred to Hyaenodon brachyrhynchus. Isolated teeth fit morphologically the material from La Débruge (Middle Ludian). However, biometric analysis suggests an Upper Ludian age. 

A new genus, Eppsillycteris, is erected for Adapisorex? allglicus COOPER, 1932, from the earliest Eocene Blackheath Beds of Abbey Wood, London, England. Various derived character states indicate that it belongs to the order Chiroptera (bats) rather than to the extinct "insectivore" family Adapisoricidae. Other derived character states are shared with fossil and modern members of the family Emballonuridae. Placement of the new genus in this family extends the record of the Emballonuridae back in time by about 10 million years. It is the earliest record of a modern bat family and one of the earliest bats. This implies that the differentiation of at least some modern bat families took place in the Palaeocene, where no authenticated records of bats yet exist. The primitive characters of the earliest bats make the family Nyctitheriidae an unlikely stem group for the order Chiroptera. A tentative plausible alternative exists in some unnamed upper molars from the Palaeocene of Walbeck, Germany. Wyollycteris chalix, described as a bat from the Late Palaeocene of Wyoming, U,S.A., fits better in the family Nyctitheriidae. 

The Squamates from Sainte-Néboule are charateristic of the Upper Eocene from the Phosphorites du Quercy. Most genera known from this locality do not survive the « Grande Coupure ». Cadurcoboa insolita gen. nov., sp. nov. is a small Boidae very characteristic of the Upper Eocene. 

Based on isolated teeth two new scyliorhinid shark species, Scyliorhinus biformis nov. sp. and Scyliorhinus suelstorfensis nov. sp., are described from the Sülstorf Beds, early-middle Chattian, of Mecklenburg, northeastern Germany. They form part of a speciose assemblage of necto-benthic sharks and batoids which populated the warm-temperate to subtropical upper shelf sea of the south-eastern North Sea Basin. 

Description of four new rodents from a recently discovered locality at Montalban. Theridomys crusafonti nov. sp. is considered as the ancestry of T. Iembronicus. Theridomys varian: nov. sp. includes «Theridomys» morphotypes and «Blainvilllimys» morphotypes; it could be ancestral to B. blainvillei. Pseudoltinomys nanus nov. sp. represents a new lineage paralleling in evolution that of P. gaillardi (which is equally found at Montalban). Pseudocricetodon montalbanensis nov. gen., nov. sp. designates a lineage of very small Cricetidae accompanying Eucricetodon. With these well defined new species and six others present in the locality, Montalban appears as the best faunal reference point within the biochronologic zone of La Sauvetat.
As an annex, discussion of two rodent specimens from the classic localíty of Tárrega, close in age to that of Montalban. 

We report the discovery of amniotic eggshells and theropod teeth from the Late Cretaccous  period in Morocco.The megaloolithid family represents the only known dinosaur egg remains in the north part of Africa and attests indirectly to the occurrence of sauropod dinosaurs (titanosaurids) in the Maastrichtian of Africa.

  

Biographie et liste des publications de S. Garimond. 

The study of all the available skulls allows us to review the systematic relationships of the longirostrine Steneosaurus from the Middle Jurassic of western Europe. Up to now, Aalenian and Bajocian deposits have not yielded any significant Steneosaurus remain. In the Bathonian, the only valid longirostrine species, S. megistorhynchus, is known in the Britain-Normandy Basin, the Poitou and the Lorraine. In the Callovian, most of the longirostrine Steneosaurus remains can be attributed to the species S. leedsi. Nevertheless, some remains from the Middle Callovian of Poitou (France) show important differences with S. leedsi. A new Steneosaurus species, only known in Poitou, is created and named S. pictaviensis. The specific characters are carried by the skull (preorbital pit well marked, orbit and ptetygoid fossae shapes), by the mandible (symphysis shape) and by the teeth (ornamentation). S. megistorhynchus is probably situated near the stem of the Callovian species but remains from the Bathonian and Lower Callovian are very scarce and it is very difficult to precise the phylogenetic relationships between the longirostrine species of the Middle Jurassic.
  

Confident attribution of bipedal tridactyl dinosaur tracks to theropods or ornithopods can be challenging. Here we describe trackways produced by tetanuran dinosaurs, previously attributed to hadrosaurs, from Coahuila State, northeastern Mexico. Multiple trackways headed in the same direction suggest gregarious behaviour in these late Campanian theropods.  

Since the 19th century, thousands of footprints were observed in the geological series of the French Mesozoic. All are located in the Triassic and Jurassic. After a promising beginning, in France, it is only a few papers which will be published in the first half 20th century, unlike the USA and of others countries of Western Europe. One ought to wait about 1950 for a revival and now they are nearly 200 papers which were devoted to the ichnofossils. The literature abundance and the renewed interest of the naturalists for the palichnologic studies decided to us to write a synthesis work. This one begins with a stratigraphic inventory in which, localisation, age and paleontological contents of about 180 fossiliferous sites are specified. After having pointed out the followed methods, the footprints paleontological interpretation is then approached in detail and the results obtained are replaced in stratigraphy to deduce the fauna evolution during the Mesozoic. So, it appears that Ichnologic data, more varied and rich in the Triassic and Liassic than those relating to the bones, very rare for the considered periods, are very informative. The middle Triassic (Anisian-Ladinian), thus reveals Cotylosauria, Lepidosauria, Crurotarsi with Rauisuchia, Ornithosuchidae, Crocodylia and Dinosauromorpha more the "Prodinosauria": Dinosamiforme whose skeletons are known in Argentina but only in Ladinian. The rather fast domination of Dinosaurs during Norian is also as well shown. The almost exclusive presence of their footprints, up to fifty cm long, in the Lower Hettangian indicates their supremacy in the environments. Footprints characterise not very deep life places located between inter-supratidal limits and often out of water. Sedimentologic and Palaeontologic studies showed that they were great coastal spaces during Middle Triassic, flood-plain with sebkhas while Upper Triassic, and a large !!coastal marsh!! in Grands-Causses during Liassic in which, mainly, fine stromatolithic layers were deposited. During the same periad, bay beaches spread in Vendée. During the Middle Jurassic, they are also brackish to lacustrine environments and recifallagoons in- the Upper Jurassic. Numerous measurements of the footprints and trackways directions showed that the animaIs moved there in weil defined directions, for long periods. They seem due to the palaeotopography of the life environments relatively stable. Also, the discovery of vegetal radicular networks and small footprints far away from the continental borderlands has suggested that the animals continuously lived in these palaeoenvironnements, belonging to large ecosystems, where the sedimentation rate was weak. This explains that thebadies could not fossilize there but only their footprints through the cyanobacterian action in main cases. From the vertical distribution of different ichnospecies, defined with adapted statistical methods, explained in this work, a palichnostratigraphy was established for the Middle Triassic. Although the footprints are also abundant in Hettango-Sinemurian of "Grands-Causses" and the Vendée, it was not possible, up to now, to establish any zonation in this series; Probably because the palichnofauna is too little diversified there, currently reduced to a majority of Theropods II-IV tridactyl traces.
  

Eosimiid and amphipithecid primates document a long and significant history of primate evolution throughout the Eocene in Southeast Asia. Despite the absence of a comprehensive post-Eocene fossil record, it was generally hypothesized that both families left no descendant in Asia. Recently, two new small-bodied taxa, Bugtipithecus and Phileosimias, have been recovered in early Oligocene coastal deposits from the Bugti Hills (Balochistan, central Pakistan) and referred to the families Amphipithecidae and Eosimiidae, respectively, on the basis of dental fossil remains. In this paper, we provide more exhaustive description, comparison, and discussion of these taxa. As for tarsiid and sivaladapid primates, the persistence of eosimiids and amphipithecids into the Oligocene clearly demonstrates that low latitudes of South Asia provided a continuous access to tropical refugia during the climatic deterioration characterizing the late Eocene-early Oligocene interval, which was seemingly lethal for primate communities elsewhere across the Holarctic continents. As a contribution to the ongoing phylogenetic debates regarding the position of eosimiids and amphipithecids on the primate family tree, we have performed a cladistic analysis in a high-level primate systematic context in order to assess the position and the role of these new taxa in that phylogenetic issue. Our results support the view according to which eosimiids and amphipithecids (and by extension Phileosimias and Bugtipithecus, respectively) are stem anthropoids. These fossils from Pakistan document an unsuspected Oligocene phase of the evolutionary history of anthropoid primates in southern Asia, which clearly enhances the extent of the anthropoid radiation in this province during the Paleogene. Several phylogenetic and paleobiogeographic aspects are discussed, notably the intra- and inter-relationships between Paleogene Asian and Afro-Arabian anthropoids, and the resulting potential dispersal models between both land-masses during the Paleogene. 

Pour la première fois un spécimen de lagomorphe a été récolté en Quercy. L'intérêt de ce fossile tient surtout à son âge géologique inattendu, qui recule considérablement la date de première apparition en Europe de cet ordre de mammifère, Ceci nous paraît justifier une nouvelle réflexion sur la « grande coupure» oligocène, 

A short review of the genera and species of snakes from the Quercy's phosphorites described by former authors is followed by the study of specimens recently collected. Most of these latter specimens belong to the Aniliidae, Boidae, Colubridae and to the Scolecophidia; the precise systematic position of some of them is not defined yet. The following genera and species are described: Eonilius europae nov. gen. and nov. sp., Platyspondylia lepta nov. gen. and nov. sp., Coluber cadurci nov. sp. and Dunnophis cadurcensis nov.  sp.; a species described by de Rochebrune (Palaeopython filholi) is revised. One of the most important conclusions of this study is that the Colubridae appear as early as the Upper Eocene. The stratigraphic repartition of these snakes shows that the rich Upper Eocene fauna is followed by the very reduced fauna of the Lower Oligocene; then the oligocene «Grande Coupure» corresponds to a very important faunistic renewal. The fauna remains poor and little diversified during the beginning of the Middle Oligocene after which there is a new faunistic explosion. 

A new hypothesis for the phylogenetic relationships of recent anomalurids and graphiurids is proposed, based on information from evolutionary lineages of Paleogene European rodents, particularly Gliridae, and Eocene Algerian Zegdoumyidae. Differences in first occurrences, in paleogeography, and in infraorbital structure in glirids (protrogomorphy and pseudomyomorphy) and graphiurids (hystricomorphy) separate Graphiuridae from Gliridae (Graphiurinae is here raised to family rank). Similar considerations, and dental morphology, suggest that Anomaluridae (appearing in the late Eocene) and Graphiuridae (appearing in the Pliocene) are related to early Eocene Zegdoumyidae. 

Le matériel provenant de nouvelles fouilles dans les phosphorites du Quercy, soumis aux divers spécialistes, a conduit à la soutenance de plusieurs thèses d'état: sur les rongeurs (Hartenberger, Vianey-Liaud), les Chiroptères et Insectivores (Sigé), les Artiodactyles (Sudre), les Squamates (Rage) et, en partie, les Chéloniens (De Broin). Chacun dans son domaine, à côté des conclusions d'ordre évolutif, paléogéographique ou paléoécologique, a pu établir des successions stratigraphiques des gisements du Quercy qui se sont révélées largement concordantes. Ainsi la succession des faunes du Quercy est actuellement l'une des mieux établies. Dans ce contexte, les différents spécialistes ont décidé de conjuguer leurs efforts dans l'étude monographique de plusieurs gisements repérés le long de cette échelle, afin de rassembler l'information paléontologique sur des faunes bien précises et de tenter d'obtenir des indications d'ordre taphonomique. 

Near "la Lieude", in the Lodève basin, more than a thousand of footprints are distributed in a twenty of trackways which amounts to 220 m length. They have been found on calcareous siltstone level in the B site named also "Réserve Naturelle Volontaire". This last is located in the Saxonian summit dated Upper Permian. "La Lieude" tracks are described by using statistical methods then they are compared with others from the world Permian. What allows to distinguish 4 following ichnotaxa: Lunaepes ollierorum nov.ichnosp., Merifontichnus thalerius nov. ichnogen. and nov. ichnosp., Planipes brachydactylus nov.ichnosp. and Brontopus circagiganteus nov. ichnosp. All these traces are attributed with possibility or probability to Therapsida or to Therosauria, except Brontopus circagiganteus nov. ichnosp. that could be due to Caseamorpha. All these animals whose sizes have been estimated between l and 5 m lived probably in a playa environment.The biological and sedimentological data from "la Lieude" footprints levels compared with informations provided by the tracks orientations, suggest the following scenario. Animals coming from the North have crossed a sandy channel bank with plants zones by directing to the South for the majority. Maybe, they were going to the lacustrine part of the playa, close to "la Lieude" footprints that they have just trampled on. 

The discovery in the Turolian (Late Miocene) of Dytiko 3 (Macedonia, Greece) of a complete skull with mandibles and cervical vertebrae, atlas and epistropheus, is a very important contribution to the knowledge of the subfarnily Chalicotheríinae. After the description, the comparison with other specimens of Miocene chalicotheres permits the revival of the generic name Macrotherium with a new species M. macedonicum. This genus is mainly characterized by a short snout and an inflated cerebral skull. It coexists during the Miocene with Chalicotherium. A cladistic analysis leads to conclusion that the species which has been described from the Early Middle Miocene of Rusinga must be identified as the type-species of a new genus: Butleria.





  

Abstract: Paleontological exploration in the Turkana Basin near Ileret, Kenya yielded the most complete adult elephant cranium (KNM-ER 63642) known from the late Miocene to mid-Pliocene. KNM-ER 63642 derives from the lower Lonyumun Mb. of the Koobi Fora Fm. and dates to the early Pliocene, >4.3 Ma. The cranium is immense in size and preserves most of its structures including left and right M2-3, permitting its comprehensive comparative study and secure taxonomic assignment to Loxodonta adaurora. Features distinctive of the species and exhibited by KNM-ER 63642 include very elongate, divergent tusk alveoli, a short, biconvex cranial roof, anterosuperior angulation of the occipital planum, non-inflated occipital planum and absence of supralateral parietal "bossing," broad, flat premaxillary nasal processes, broad, laterally downturned nasal aperture superior to the level of the orbits, and M3s with wide, subhypsodont plates that are parallel-faced and separated by U-shaped transverse valleys. The M3s also exhibit characteristic L. adaurora traits of greatest width at their bases, rounded cross-sectional shape, thick enamel, abundant cementum, and strong anterior and posterior accessory conules. Of extant taxa, KNM-ER 63642 most closely resembles crania of African elephants. Its inclusion in the Loxodonta clade is tenuous, however, because shared features are either symplesiomorphic or are difficult to test for synapomorphy due to the poor fossil record of crania of late Miocene-early Pliocene elephants. Overall, the cranial morphology of KNM-ER 63642 is unexpectedly advanced for an elephant of its antiquity. Its anteroposterior compression and height are concordant with efficient proal masticatory action, indicating that by the early Pliocene L. adaurora evolved craniodental adaptations in phase with feeding preference for C4 grasses. The advantage of synchrony of morphology and behavior is reflected by the dominance of the species in the greater Turkana Basin during that interval.