A systematic analysis of an hipparionine horse assemblage from Maragheh, Iran is made. A brief orientation to systematic philosophy and informal superspecific characterizations of some Old World hipparionines is given as a background to this work. A character state analysis of skulls is made, and has revealed five distinct species. A character state and stratigraphic trend analysis of isolated check tooth and postcranial remains, with known provenance, is also made. These two combined analyses reveal that the most resolute discrimination of hipparionine species and their evolutionary relationships occurs when multiple character complexes of associated skulls, maxillary and mandibular dentitions are made. When this is not possible, skulls have provided the best basis for discriminating species and their evolutionary relationships. Traditional characters of isolated cheek teeth and postcranial remains are shown here to offer limited information content for hipparionine phylogenetic systematics. The systematic portion of this study includes a comprehensive description of cranial and postcranial remains, and has further corroborated the distinction of five species which belong to at least three superspecific groups including: «Hipparion» geltyi sp. nov., Group 1; Hipparion prostylum (s. l.), and Hipparion campbelli sp. nov., Group 3; «Hipparíon» aff. moldavicum and «Hipparion» ?matthewi, Group 2. These species stratigraphic ranges and evolutionary relationships are also given here and argued to be important for establishing future hipparionine geochronologic correlations between a number of Eurasian late Miocene provinces. 

Murids and gerbillids (Rodentia, Mammalia) recovered from four Upper Siwalik localities; Moginand (around 3.5-4.5 m.y.), Kanthro (around 2.5 m.y.), Ghaggar (around 2 m.y.) and Nadah (around 1.8-2 m.y.) are described herein. A comparison of Mus linnaeusi sp. nov. reveals its close relationship towards the house mouse Mus musculus. In contradiction to earlier proposals that Golunda migrated to the Indian subcontinent from Africa sometime during Late Pliocene, it is suggested here that Golunda (in the form of Golunda tatroticus sp. nov.) evolved from Parapelomys robertsí of Late Miocene deposits of Siwaliks. Golunda tatroticus sp. nov. exhibits a progressive relationship to the extant Indian Bush Rat Golunda ellioti through Golunda kelleri and Golunda sp. Tatera pinjoricus sp. nov. is considered here to be a link between the extinct Abudhabia kabulense and the extant Tatera indica.

  

Abstract: Paleontological exploration in the Turkana Basin near Ileret, Kenya yielded the most complete adult elephant cranium (KNM-ER 63642) known from the late Miocene to mid-Pliocene. KNM-ER 63642 derives from the lower Lonyumun Mb. of the Koobi Fora Fm. and dates to the early Pliocene, >4.3 Ma. The cranium is immense in size and preserves most of its structures including left and right M2-3, permitting its comprehensive comparative study and secure taxonomic assignment to Loxodonta adaurora. Features distinctive of the species and exhibited by KNM-ER 63642 include very elongate, divergent tusk alveoli, a short, biconvex cranial roof, anterosuperior angulation of the occipital planum, non-inflated occipital planum and absence of supralateral parietal "bossing," broad, flat premaxillary nasal processes, broad, laterally downturned nasal aperture superior to the level of the orbits, and M3s with wide, subhypsodont plates that are parallel-faced and separated by U-shaped transverse valleys. The M3s also exhibit characteristic L. adaurora traits of greatest width at their bases, rounded cross-sectional shape, thick enamel, abundant cementum, and strong anterior and posterior accessory conules. Of extant taxa, KNM-ER 63642 most closely resembles crania of African elephants. Its inclusion in the Loxodonta clade is tenuous, however, because shared features are either symplesiomorphic or are difficult to test for synapomorphy due to the poor fossil record of crania of late Miocene-early Pliocene elephants. Overall, the cranial morphology of KNM-ER 63642 is unexpectedly advanced for an elephant of its antiquity. Its anteroposterior compression and height are concordant with efficient proal masticatory action, indicating that by the early Pliocene L. adaurora evolved craniodental adaptations in phase with feeding preference for C4 grasses. The advantage of synchrony of morphology and behavior is reflected by the dominance of the species in the greater Turkana Basin during that interval.
  

In dilambdodont molars the primitive crest between paracone and metacone (centrocrista) is represented by a pair of crests that join the mesostyle (postparacrista, premetacrista). The cutting action of these crests against the crests of the hypoconid is described. Dilambdodonty is a derived adaptation for greater cutting efficiency. It has evolved several times and in more than one way. 

The complex taxonomic history of the paramyid rodent genus Ailuravus is reviewed. It has been described as Hyracotherium, as a creodont carnivore and as a lemuroid primate - errors at the ordínal level that are most unusual for a rodent. The genus is a member of the poorly known subfamily Ailuravinae, probably derived from some European Early Eocene species of Paramys. Aíluravus was a large arboreal paramyid with highly rugose cheek teeth, very well developed hypocone, and a remarkably weak lower incisor. It was tropical to subtropical. Three named species are recognized, A. macrurus from the Lutetian of Messel; the genotype, A. picteti, from Egerkingen, Buchsweiler and the Geiseltal, slightly later in the Lutetian; and A. stehlinschaubi, new name, from the Bartonian of Mormont-Eclépens and Robiac. One or more unnamed species are present in the Ypresian of Cuis. The species are close to a phyletic sequence. No later representatives of the genus are known. The late Eocene to earliest Oligocene North American paramyid Mytonomys, whose relationships have been obscure, is tentatively referred to the Ailuravinae. 

Is hereby described an impressive lchnoiauna belonging to the Lower to Middle Ludian of the Gard (S. France). The slab, already cleaned over a length of 18 m, is located near the top of the Potamides aporoschema lacustrine limestone (Lower Ludian, Euzet zone). It is therefore older than the Célas sandstone deposit, and still more than the Melanoides albigensis and M. acutus marly limestone corresponding to the Upper Levels of the Ludian stage. Although biostratigraphically older than the La Débruge and Montmartre zone, the biotope shows already a sampling of very tall Artiodactyles, Perissodactyles and Carnivorous. One of the most « majestic ›› Artiodactyles, Anopolotheriipus lavocati, nov., points out a huge size type. To mention also among the Ichnotypes described, 10, the big Perissodactyle Palaeotheriipus similimedius, nov., and the big Carnivorous Hyaenodontipus praedator, nov. 

Sirenians are among the common marine fossil remains found in the Faluns deposits of western France. We describe new material of a Dugongidae sirenian from the middle Miocene Savignean facies of La Morfassière quarry (Indre-et-Loire, northwestern France) that includes a well preserved and almost complete skull associated with its mandible, several vertebrae and ribs. The cranial remains exhibit features that allow to attribute them to Metaxytherium medium, a species recorded from the middle and early late Miocene of European and Mediterranean coasts. The discovery of an associated skull and mandible of this species is unusual in this area and deserves to be reported, mostly because its preservation contributes to a better knowledge of the variable structure of its skull and teeth anatomy. For the first time the body size and weight of M. medium can be assessed through regression equations based on skull measurements. The particularly good condition of the material can be explained by the calm and deep marine environment in which it was deposited.
  

The fossil mammals collected from the Eocene of Hammada du Dra (northwest Sahara. Algeria) and two fragmentary teeth from the Lutetian of M'Bodione Dadere (Senegal) are described.
The fossils from the northwest Sahara come from a lacustrian deposit dated by charophytes (Raskyella aff. pecki, Raskyella n. sp.. Maedleriella lavocati, Maedleriella sp. et ? Peckichara sp.) as Middle Eocene or perhaps Lower Eocene (Gevin, Feist and Mongereau, 1974). Several hyracoids (3 or 4) identified from this formation extends the age of the family Pliohyracidae Osborn in Africa. Three forms appear to belong in the genera Megalohyrax, Titanohyrax and perhaps Bunohyrax which have been know until now only from the lower Oligocene of the Fayum (M. gevini n. sp. ; T. mongereaui n. sp.. ? Bunohyrax or Megalohyrax indet.). Another hyracoidof small size is referred to a new genus, Microhyrax (M. lavocati n. sp.).
Helioseus insolitus n. g. n. sp. is described without ordinal assignment. Azibius (Sudre, 1975) which has been the subject of questions and interpretations is reviewed.
Only one tooth from the Lutetian of M'Bodione Dadere is complete enough to interpret. lt probably belongs to a condylarth and demonstrates for the first time, the presence of the order in Africa. The second tooth is too fragmentary for comment.
In conclusion., the paleobiogeographic role of Africa at the end of the cretaceous and the beginning of the Cenozoic is discussed. 

A fragmentary mandible and maxilla of a small sized Agriotherium of a young individual discovered from a Pliocene fissure filling in Xiaoxian county (Anhuei Province, China) are described. Judging from the morphology of the dentition and its dimensions the new material can be identified as Agriotherium inlermedium (STACH l957). Hendey's proposition (1980) that the Agriotherium species are derived from Indarctos is reconsídered on the basis of the new documents. As a result of a more general phylogenetic discussion it can be stated, that: 1. the supposed size increase as well as other trends, leading from Indarctos to Agriotherium are untenable ; 2. there are no positive indications to assume a phylogenetic transition of these two genera. 3. there are no real arguments in favor of an adaptational reversal in the evolution of Agriotherium. Hence, many features of that genus supposed by Hendey to be derived are plesiomorphic ; 4. regardless of the previous points it is methodologícally impossible to establish direct ancestor - descendant relationships between Indarctos and Agriotherium species, as Hendey did. Based on the data available and especially on the characters of the new material from China it is more likely that Agriotherium and Indarctos are two genera which developed independently. While advanced Agriotherium species, e.g. A. africanum, resemble in some respects Indarctos by adaptational analogies, more primitive species, e.g. Agriotherium intermedium, are quite dissimilar to lndarctos. While Indarctos might be derived from an Ursavus like forerunner, Agriotherium has its roots more likely somewhere in between Ursavus and the Hemicyon-group. 

Recent collecting of fossil vertebrate remains from the lowermost member of the Aquia Formation (Paleocene), has enabled me to report here for the very fIrst time, the earliest occurrence for the teeth of Palaeocarcharodon in the fossil record of the New World.
This report represents only one species of neoselachian from this locality, the remaining fauna of which will subsequently be described. 

Some new material of percrocutas from the late Miocene of Axios valley (Macedonia, Greece) is studied. They have been found in the locality of "Pentalophos 1" (PNT). The material has been described and compared with the known late Miocene percrocutas of Eurasia. This comparison indicates that it can be identified as Dinocrocuta gigantea (SCHLOSSER, 1903). A maxilla of a percrocuta, named ”Hyaena" salonicae, was found in the same area (Andrews, 1918). "Hyaena" salonicae is smaller than the PNT material. It is also compared with other material from Eurasia while its taxonomic and age problems are discussed. It belongs to Dinocrocuta and shows close relationships with D. robusta and D. senyureki; its age can be considered as late Vallesian-early Turolian. The age of the locality PNT is also discussed and a possible Vallesian age is proposed for it. 

Prolagus sardus is the last representative of the diverse lineages of European endemic ochotonids. It is also the most abundant in the collections. The previous studies made of this species have established rather well its dental morphology, its phylogenetic position, its geographic and temporal distribution, and its intraspecific individual variation. On the other hand, no osteologic study has fully utilized the superb material from Corsica and Sardinia  collected by Forsyth Major.
Nearly all of the parts of the skeleton are represented in this material; they are here described and figured. Comparisons are made with Ochotona, the only surviving genus of the family, as well as with the living leporids, in particular Romerolagus, Oryctolagus and Lepus. A brief examination of the dentition reveals some particular characters of the incisors. The osteologic study allows limited interpretations to be advanced concerning the posture, mastication. locomotion and some other adaptive features of P. sardus.
P. sardus appears as an incontestable ochotonid, but it differs from Ochotona by some characters which are found in leporids. Certain of the characters are here judged primitive for lagomorphs. The genus Ochotona therefore can no longer be considered as the image of a primitive lagomorph, because in spite of the lack of cursorial adaptations which differentiate it from leporids, it presents other specializations which are common to it alone. 

Designated the type-locality of a late Eocene paleomammal zone, Robiac has recently been the object of important excavations. The first results of the new collecting, as  well as a revision of the material in old collections, are given in this work.
Two stratigraphic section, cutting through the two sites presently distinguished (Robiac-Nord and Robiac-Sud) reveal the lithologic variation at the base of the Fons Limestone and the localization of three fossiliferous beds at Robiac-Sud.
The molluscan fauna and the flora (charophytes), which have already been described, as well as that of the lower vertebrates, have been listed.
A list of 46 mammalian species (only 16 species were known previous to 1964) has been established. The micro-mammals, nearly all new in this fauna (marsupials, insectivores, bats, rodents, primates, and some smallsized artiodactyls), Were obtained only after screen-washing of the matrix; about 4 tons of sediment were thus treated.
The artiodactyls have been the most extensively analyzed; 6 genera, of which one is new, have been recognized. The latter is described as Robiacina minum n.g., n.sp., and represents a very small artiodactyl of the family Anoplotheriidae. The taxonomie status of certain species formerly described has been clarified by the designation of lectotypes (Cebochoerus robiucensis, Catodonerium robiacense, Xiphodon castrense).
The paleontologic corrélations at present possible between the late Eocene faunas have allowed the relative positions of te different French localities of this age to be established; the Guépelle locality, it seems, could define in the future a new paleomammal zone.
  

Extensive sampling of the Siwalik deposits of the Potwar Plateau of northem Pakistan and from the Zinda Pir dome of central Pakistan has produced a fossil record of Miocene ctenodactylids that can be correlated with the paleomagnetic time scale. The early Miocene Prosayimys flynni (n. gen., n. sp.) is recognized as the first ctenodactylid in the Indian subcontinent. Prosayimys is ancestral to Sayimys. From the late early Miocene to the early late Miocene, there is an anagenetic succession of three species of Sayimys: S. cf. S. intermedius, S. sivalensis, and S. chinjiensis (n. sp.). Sayimys chinjiensis gave rise to the late late Miocene S. perplexus. A second lineage is represented by Sayimys minor, S. sp. A, and S. sp. B. 

A new Devonian vertebrste locality has been discovered in 1976 in the basal part of the Khush-Yeilagh Formation in the eastern Alborz Mountains of Iran. The fossils occur in a band one centimeter thick which is identifiable at other outcrops in the area. A preliminary study of the remains has yielded the following faunal list : Placodermata (Phlyctaeniida indet.. Groenlandaspididae indet.. Coccosteidae indet., Holonema sp., Ptyctodontida indet., Antiarcha indet., Bothriolepis cf. kwangtungensis), Elasmobranchii (Ctenacanthidae indet.), Acanthodii (Gyracanthus sp., « Onchus » overathensis, Ischnacanthiforme indet.), Dipnoi (? Dipteridae indet.), « Crossopterygii ›› (Onychodus cf. sigmoides, Holoptychiidae indet., Osteolepididae indet.). This fauna is older than the Middle Givetian and probably of Middle or Lower Eifelian age. Owing to its diversity, it may prove possible to use it in the future as a reference fauna for the study of the Devonian vertebrates in Central Asia and the Middle East. 

Fossils of the Trionychidae, Bataguridae or Emydidae, Palaeophis and Crocodylia from Chattahoochee, NW Florida, USA, represent the first report of an Eocene reptile fauna from Florida. 

Herein we describe a small but relatively diverse assemblage of fossil fishes derived from the lower Oligocene (Rupelian) Red Bluff Clay at site AMo-9 in Monroe County, Alabama, USA. Identified amongst the remains are 15 unequivocal taxa representing 11 families within five orders, and one additional taxon represents an unknown order and family. Taxa identified include Eostegostoma sp., Otodus (Carcharocles) sp., Mitsukurinidae/Carchariidae indet., Macrorhizodus praecursor, Galeorhinus sp., Negaprion gilmorei, Physogaleus sp., “Sphyrna” sp., Galeocerdo sp., cf. “Aetobatus” sp., Sphyraena sp., Xiphiorhynchus kimblalocki, Xiphiorhynchus sp., Cylindracanthus ornatus, and C. rectus. Several additional fossils could not be identified beyond Lamniformes, Carcharhiniformes, and Teleostei, but they likely belong to one of the identified taxa within this paleofauna. All of the fishes previously reported from the Red Bluff Clay within the entirety of the Gulf Coastal Plain of the USA are otolith-based, and each of the 15 unequivocal taxa reported herein are important new records for this lithostratigraphic unit. In particular, the Eostegostoma sp. and Xiphiorhynchus spp. specimens represent the first occurrences of these taxa in Alabama. The specimens of C. ornatus, Eostegostoma sp., and X. kimblalocki are stratigraphic and temporal range extensions from the middle and late Eocene into the Rupelian Stage of the Oligocene. Other described taxa may represent transitional forms between those described from the late Eocene and late Oligocene within the region. This study provides a tantalizing preliminary view into faunal transitions that occurred amongst marine fishes across the Eocene/Oligocene boundary within the Gulf Coastal Plain of the USA. 

Valverde's cenogram method is used to analyse mammalian communities from the late Eocene to late Oligocene of southern France, mainly from the "Phosphorites du Quercy". Cenogram analysis involves plotting the size of each component species in a fauna on a semilog diagram in rank order, permitting fossil faunas to be compared with Recent ones. The configurations of Recent communities serve as models for establishing the general environmental characteristics of fossil mammalian faunas. This method of analysis applied to faunal sequence can reveal major and sudden ecological perturbations. The paleobiogeographical event (i.e. the mammalian immigration wave) at the Eocene-Oligocene Boundary in western Europe, known as the «Grande Coupure", is here shown to represent a drastic and sudden ecological change: late Eocene tropical environments in Europe deteriorated rapidly turning to subdesert or desert environments al the beginning of the Oligoccne. 

Re-evaluation of the small collection of mammal fossils from the Changxindian Formation near Beijing, China indicates the following taxa are present: Eutheria, Hypsimilus beifingensis, cf. Miacis sp., Anthracotheriidae and Forstercooperia grandis. The presence of Forstercooperia grandis indicates an Irdinmanhan age and does not support previous assignment of a Sharamurunian age to the Changxindian Formation. 

The dentition of selachians is characterized by an often very pronounced heterodonty involving a great morphological diversity. Despite this fact, the dentitions of selachians can be grouped in a rather reduced number of dental types corresponding to trophic adaptations: grasping, tearing, cutting, crushing, grinding and grasping-grinding type. The numerous exemples of convergence and parallelism that can be observed in fossil selachians and between Recent and fossil ones is the result of this reduced number of dental types. These dental specialisations allow to try a reconstruction of the way of life of fossil forms.