The analysis of oxygene isotope variations as well as paleobotanical data suggest that the Oligocene/Miocene boundary corresponds to a transitional period marked by floristical and climatic variations. During this period, the pyreneo-alpine tectonics has contribued to modify the geography and western Europe landscapes. Faunal changes (appearances, extinctions, migrations) are observed in different mammalian groups, notably in the rodents. A study of the evolutionary trends and patterns in paleogene rodents is involved for the period ranging from level MP 28 of the Late Oligocene to the Early Miocene, including the Oligo-Miocene boundary.
The Rodents fauna from the sites of Venelles (Bouches-du-Rhône District, France) and Thezels (Lot, France), previously mentionned in litterature, have been studied. The first description of the Eomyidae of La Milloque (MP 29) has been completed. These faunas are compared to those from various localities dating from the considered period. In La Milloque, a new representative of the Eomys species is described next to a form close to Rhodanomys hugueneyae ENGESSER, 1987. It is the Eomys milloquensis nov. sp., the likely descendant of Eomys quercyi COMTE & VIANEY-LIAUD, 1987. Two new species are also described in Thezels: Eucricetodon thezelensis nov. sp., resulting from a likely and local evolution of Eucricetodon praecursor (SCHAUB, 1925) from La Milloque, which, in the same geographic area, could be at the origin of Eucricetodon hesperius ENGESSER, 1985 from Paulhiac. Plesiosminthus admyarion nov. sp., quite distinct from Plesiosminthus schaubi VIRET, 1926, which announces Plesiosminthus myarion SCHAUB 1930. Venelles 'Plesiosminthus schaubi population is considered as a sub-species, named Plesiosminthus schaubi meridionalis nov. subsp. New phylogenetic patterns are proposed. Among the Eomyidae, a quantification of various features of the M1-2/ crown (hypsodonty, degree of abrasion, occlusal angle, state of development of the I and V anticlines), and a comparison with the occlusal diagram of the other teeth among various other populations allows a more efficient separation of Eomys and Rhodanomys genera. In Western Europe, and within this period, it finally does not seem possible to gradually connect the genus Eomys to the genus Rhodanomys. The evolution of the Eomys quercyi - milloquensis lineage seems to underline a similar evolution to that which may have led from the Eomys to the Rhodanomys form. The latter which appears totally accomplished at level MP 29 of the Oligocene is considered as an immigrant. If we compare the most representative species of the Venelles, Thezels, and Coderet sites, (i.e. Rhodanomys, Eucricetodon, Adelomyarion, Peridyromys, Plesíosminthus), it becomes impossible to confirm their biochronological separation. The noticeable differences between the populations may be interpreted as geographical variations. An explanation to these variations, and to fauna's evolution during the Late Oligocene and Early Miocene can be found in the environmental modifications, supported by isotopic, paleobotanical and sedimentologic analysis. A tentative reconstruction of the environments is attempted by the cenogram method. The analysis of the fluctuations of fauna's diversity shows variations which may be correlated to a drop in temperature at MP 29, during the Late Oligocene, followed by an increase in temperature along with an aridity phenomenom, during the basal Miocene (MN O).The confrontation of various methods give the opportunity of reconstituting and comparing the evolution of the environment of three sequences of sites chosen from different regions. Ecological affinities of various rodents' species are being examined. It is possible to consider that the integration of all the conclusions resulting from this study should lead to an explanation to the evolution of rodents for the period around the Oligocene-Miocene boundary. The site of Coderet- level 3- would be posterior to the latter, at the beginnig of the Miocene, and would mark the level MN 0 of the Aquitanian.

  

The Caudata are known by two Salamandridae ; one of them is attributed to the genus Megalotriton. The Pelobatidae form the major part of the Anura ; a few bones indicate also the presence of Neobatrachia. 

 A new genus of hystricognath rodent with two new species, Soriamys gaimanensis and Soriamys ganganensis, from the Colhuehuapian Age (Early Miocene), Sarmiento Formation, of Patagonia is described. The first species comes from the south barranca of the valley of Rio Chubut, near Gaiman locality (Chubut Province). It is known through a great number of dental remains in different stages of wear. The teeth are protohypsodont, with cement in the principal valleys. P4 are simpler, more oval inoutline and without a differentiated hypocone-hypoflexus. Upper molars are pentalophodont in early stages of wear and become bilobated in middle and advanced stages; M3 have a temporary third posterior lobe due to the maintenance of the braquiflexus on the posterior wall of the tooth. Lower teeth have a completely asymrnetrical pattern in relation to the uppers. The molars are trilophodont, with only one complex crest in the trigonid composed of the fusion of the anterolophid and mesolophid. The dp4 have a derived pentalophodont pattern due to the loss or fusion of the mesolophid and the development of an accessory transverse crest between the anterolophid and metalophid. The second species is known through a complete skull and jaw and other skull fragments with the dentition, proceeding from Pampa de Gan Gan, Chubut province. Respect to the first species, the teeth are higher crowned, with an earlier occlusal simplification and a third posterior lobe of M3 more developed and persistent. The skull and jaws show many chinchilloid characters, such as jugal bone with an ascending process, concave palate, very prominent dorsal shelf of the masseteric fossa, nasolachrymal duct opened laterally on the maxillary and very developed lateral mandibular fossa. Moreover, Soriamys is closely related to Cephalomys by sharing a similar asymmetric dental pattern and other dental and mandibular characters. These two genera constitute a natural group, the Cephalomyidae, with peculiar dental characters, like the asymmetric pattem of the upper and lower teeth; Cephalomyopsis, and probably Litadontomys, can be considered part of this group. Scotamys and Perimys constitute a distinct lineage (the Neopiblemidae or Perimyidae), more closely related to the chinchillids. Many characters shared by cephalomyids, eocardiids and caviids suggest a closer affinity between the Chinchilloidea and the Cavioiidea as a whole. Abrocoma shows also an asymmetric dental pattern and other chinchilloid features that suggest some degree of relationship with the cephalomyids, still not well determined. 

Three new fossiliferous localities, two of karstic origin, Castelnou 3 and Font Estramar, respectively Late Upper Miocene and Lower Pliocene, and one of lacustrine origin, Thuir, Lower Pliocene, add data about the transition between Miocene and Pliocene faunas of rodents in southern France. An unexpected association of taxa was present in the late Upper Miocene, including between others, Myocricetodon, Hispanomys, Ruscinomys, Cricetus barrierei, Promimomys and a new species of Stephanomys, S. dubari nov. sp. Myocricetodon is still known in the Lower Pliocene. It is shown that the large field-mice known since the Late Upper Miocene belong to two different lineages, on one side, A. jeanteti, on the other side, A. gudrunae followed by A. gorafensis. Biochronological and biogeographical implications are discussed. 

The presence at Silveirinha of one of the earliest, ? Late Paleocene or Lowermost Eocene, european representatives of the genus Diplocynodon is based mostly on isolated bones and teeth (often from juveniles). This small-sized form is the only crocodilian so far recognized in this site. The longevity of Diplocynodon in Portugal becomes much extended; the genus survived there until the Middle Miocene at least. A discussion on the possible affinities with other eocene Díplocynodon and especially those from Cubillos-Valdegallina (Zamora, Spain) is presented. On the other hand, differences have been detected in comparison with: Díplocynodon tormis, from the middle Eocene of the Douro basin in Spain, which may belong to another phyletic line; and the aff. Diplocynodon from Dormaal (Belgium) and Le Quesnoy (France), nearly contemporaneous of Silveirinha. The Silveirinha Diplocynodon and many other data strongly suggest moist, subtropical, quite limited in space environments related to an alluvial plain crossed by small, meandering channels. 

The first part of this study was devoted to a descriptive analysis of teeth morphologies among the vespertilionine bats. This leads now to a tentative synthesis, providing views on the systematics of the group. The results could be seen according to three distinct but closely related purposes : 1 - the sorting of the genera contents in order to conform the genera units to homogeneous taxa that could represent natural issues of evolutionary lineages ; 2 - the investigation of relationships between extant genera in order to infer the possibilities of common origin ; 3 - according to the preceeding items and to the observed evolutionary trends, a tentative phylogeny, modest and cautious. The contents of many genera are sorted : Leuconoe is removed from subgeneric to generic position, whereas Myotis becomes a subgenus of it ; the myotodont species are cleared away from the Pipistrellus genus ; Glischropus and Scotozous are synonymized within Pipistrellus ; Hypsugo is raised to the generic level ; some species previously ranged within Pipistrellus will form provisionally a collective group, Attalepharca nov. ; the Eptesicus genus is broken up, the excluded species being grouped within Nycterikaupius gen. nov. ; the Nycticeini tribe is defined again after exclusion of Otonycteris , Scotoecus, Scotophilus , and addition of Hesperoptenus ; the species la io and Pipistrellus tasmaniensis are removed to Eptesicus (n.s.) and Pipistrellus dormeri to Scotoecus. Groupings of genera are stated according to the main evolutionary trends of I1/. The relevance of these is often warranted by close morphologic similarities of other teeth. This leads to a recognition of the major evolutionary radiations which occurred in the group. The filiations schematized at the end of the work show the dental relationships observed between the extant genera, and could represent a phylogenic framework. Two major facts are to be underlined : 1- the early divergence of leuconoids ; 2 - the successives crossings to myotodonty from the nyctaloid flow. Fossil data from the literature are punctually and tentatively incorporated within phylogenic sketches. 

A late Pleistocene deposit of 60 species of vertebrates and 12 of invertebrates is described from Robinson Cave, Overton County, Tennessee, U.S.A. Forty-eight species of mammals are represented by at least 2,483 individuals; 10 % are extinct, 10 % occur in the state only as boreal relicts in the Great Smoky Mountains; 23 % no longer occur as far south as Tennessee; 57 % occur at or near the site today. Nínety-one percent of the Recent mammal species can be found living today in the Minnesota-Wisconsin area, approximately 10 degrees farther north. Fluorine analysis suggests a long period of accumulation. The following 10 mammalian species are recorded from Tennessee for the first time. Sorex arcticus, Microsorex hoyi, Citellus tridecemlineatus, Clethrionomys gapperi, Microtus pennsylvanicus, Synaptomys cooperi, Synaptomys borealis, Zapus nudsonius, Napaeozapus insignis, Martes americana. Six additional species are present as boreal relicts in the Great Smoky Mountains of eastern Tennessee but not at the site today : Sorex cinereus, Sorex dispar, Sorex palustris, Parascalops breweri, Glaucomys sabrinus, Mustela nivalis. Six forms are extinct: Canis dirus, Ursus americanus amplidens, Sangamona furtiva, Dasypus bellus, Mammut americanus,Megalonyx jeffersoni. Twenty-six additional species of mammals, all of the snails, birds, reptiles, and amphibians recovered from the fauna still inhabit the area today: The fauna is indicative of a cold-temperate climatic episode associated with the Wisconsin glaciation, but may be chronologically mixed. 

The family Didelphidae is represented by three species in the Sainte-Néboule site, phosphorites of Quercy (lower Oligocene, San Cugat's nivel): Amphiperatherium minutum (Aymard), Amphiperatherium sp. and Peratherium cuvieri (Fischer). Only the first  and  third species are abundant (88 and 97 pieces). This two populations are described. The marsupial fauna of the european lower Oligocene is not recognized in its entirety in this site. 

A recent campaign of excavations (1965-68) undertaken by the Laboratoire de Paléontologie of Montpellier in pockets of the Quercy phosphorites, has permitted the dating of several localities thanks to the analysis of their micromammalian fauna.
The rodents of localities belonging to the middle Oligocene (La Sauvetat zone) are the object of this study. This rodent fauna has been compared to the forms coming from three stratified localities belonging to the biochronologic zone of La Sauvetat :  Balm, Montalban and Lovagny.
The abundance of material, notably of theridomyids and cricetids, permitted a summary statistical study to be effected and gave some precise details on the variability of the species (Theridomys varians in particular). Other details were provided on certain groups, notably on the upper teeth of Scíuromys cayluxi and on the milk teeth of theridomyids. Some specimens of «Sciurus» sp. and of Plesispermophilus angustidens were collected. By this means, it has been possible to establish, for the first time, a precise upper limit to the epoch of their appearance in Europe. 

The middle Palaeocene of São José de ltaboraí (State of Rio de Janeiro, Brazil) has produced a rich and diverse fauna of boid snakes. It comprises six or seven species: Hechtophis austrinus gen. et sp. nov., Corallus priscus sp. nov., Waincophís pressulus sp. nov., Waincophis cameratus sp. nov.,"Boinae A", and "Boinae B". Moreover, two dentaries might pertain to either H. austrinus or "Boinae B", or even represent a distinct taxon. Hechtophis austrinus is assigned, with reservation, to the Erycinae. All other taxa are referred to the Boinae. The vertebrae of all taxa have paracotylar foramina, which raises the problem of the apomorphic or plesiomorphic nature of this feature. This fauna also raises the question of the presence of extinct erycine boids in South America, but it does not allow this question to be settled. 

The rodents (Cricetidae, Gliridae, Sciuridae) found in lacustrine, brackish marine and karstic sediments of Miocene age in Languedoc, assign the position of the different localities in the scale of "niveaux repères" used by mammalogists. Some detailed stratigraphical studies bring several correlations between this continental biochronological scale and the marine scale ; the most important results are the Aquitanian age of the "niveaux repères" of Coderet and Paulhiac, the Burdigalian age of Laugnac, Estrepouy, Vieux-Collonges, La Romieu and Sansan and the Langhian or Lower Serravallian age of La Grive M. The correlations between the Tethys and the Central Paratethys for the Lower Neogene profit also of these results, since the locality of Neudorf Spalte 1, 2 (Czechoslovakia) is shown to be younger than Sansan (France). The paleontological study has also several geological inferences for the Miocene of Languedoc ; with the calibration of this Miocene, we know quite precisely that the Lower Miocene is chiefly a time lacustrine sedimentation, and also that the marine Miocene sedimentation ends early in the Miocene Period, in Langhian or lower Serravallian times. 

The genus Pachynolophus, one of the poorest known of the Palaeotheriidae, includes the brachyodont forms with reduced and non-molariform premolars and with upper molars lacking a mesostyle. Quantitative characters (divers surface indications and elongation of the teeth), while demonstrating a close relationship to Hyracotherium, permit a better differentiation of the genus, confirm its specific splitting, and permit the distinction of three lineages. The skull from Memerlein is taken as the type of a new species, P. Iavocati, of which the dentition is extremely characterized by its lophiodonty, the strong reduction of the premolars and the reduction of the cingula. This characterization testifies to a late age which extends the existence of the genus quite near to the Eocene-Oligocene limit. Compared with the only two skulls known of related species (Hyracotheríum vulpiceps and Pachynolophus Iivinierensis), that from Memerlein is distinguished by progressive characters affecting diferent regions but most particularly the braincase; it is not possible, however, to isolate within this evolution the part which leads to a systematic differentiation. Modernization is translated by a considerable increase in size of the braincase, principally in the frontal region, a development of the facial region with anterior displacement of the dental series and a greater specialization of the masticatory apparatus. This evolution parallels the history of the Equidae of the North American early Tertiary, but certain particularities, the form of the alisphenoid, the presence of an anterior frontal foramen, and the structure of the paroccipital apophysis, testifies to the independance of the European forms. 

We present here a new updated counting of the perissodactyls of Robiac, the type locality of the MP 16 level of the biochronological scale of paleogene mammals and that of the Robiacian stage of Eocene Land Mammals Ages in Western Europe.
The outcrop of Robiac consists actually of two 500m apart loci, Robiac-Nord and Robiac-Sud, considered of the same age according to the current discriminating power, and is dated from -38,7 MA after the last faunal, magnetostratigraphic and climatic calibrations.
It has yielded a very abundant and rich of 21 taxa perissodactyl fauna, topped by the giant Lophiodon lautricense, last representative of the family Lophiodontidae, of which it is the last proved deposit. The Palaeotheriidae are much diversified with 5 genera and 9 species of "Pachynolophinae", 3 genera and 10 species of Palaeotheriinae. Nine taxa have been defined from Robiac: Chasmotherium depereti n. sp., Palaeotherium castrense robiacense Franzen, 1968, the genus Leptolophus Remy, 1965 with the species L. stehlini Remy, 1965 and L. magnus Remy, 1998, Anchilophus (Paranchilophus) jeanteti Remy, 2012, Metanchilophus chaubeti Remy, 2012, Lophiotherium robiacense Depéret, 1917 and Pachynolophus gaytei n. sp.
The faunal Robiac cenogram with the associated flora testify to a hot, wet and forestal environment, likely corresponding to a short warming climatic phase; the broken up fossil bones should have been carried away and then gathered in swamp areas along the banks of a meandering river.
The swarm of mammals of Robiac, the richest of contemporaneous deposits, has been followed by a drastic drop in perissodactyl diversity at the MP 17A level; a crisis which could have originated in a renewal of the global Eocene cooling. Fons 4, the type-locality of this level, is largely scarcer in perissodactyls and its cenogram testifies to a less diversified fauna, with on the whole smaller species, that likely means a cooler and drier climatic environment; a new perissodactyl diversification occurred but later.
  

A short review of the genera and species of snakes from the Quercy's phosphorites described by former authors is followed by the study of specimens recently collected. Most of these latter specimens belong to the Aniliidae, Boidae, Colubridae and to the Scolecophidia; the precise systematic position of some of them is not defined yet. The following genera and species are described: Eonilius europae nov. gen. and nov. sp., Platyspondylia lepta nov. gen. and nov. sp., Coluber cadurci nov. sp. and Dunnophis cadurcensis nov.  sp.; a species described by de Rochebrune (Palaeopython filholi) is revised. One of the most important conclusions of this study is that the Colubridae appear as early as the Upper Eocene. The stratigraphic repartition of these snakes shows that the rich Upper Eocene fauna is followed by the very reduced fauna of the Lower Oligocene; then the oligocene «Grande Coupure» corresponds to a very important faunistic renewal. The fauna remains poor and little diversified during the beginning of the Middle Oligocene after which there is a new faunistic explosion. 

Previous chronological arrangements of South American Quaternary land mammal faunas are appraised on the basis of current geological and paleontological data. Three South American late Pliocene-Pleistocene land mammal ages are conventionally recognized, from oldest to youngest, the Uquian, Ensenadan, and Lujanian ; all are defined on Argentine faunas.

     The Uquian is based fundamentally and historically on the fauna from the Uquía Formation in Jujuy Province, northwestern Argentina. Important known formations in Argentina yielding Uquian Age faunas include the sub-surface Puelche Formation (or Puelchense) near the city of Buenos Aires, and the Barranca de Los Lobos and Vorohué Formations between Mar del Plata and Miramar, Buenos Aires Province. A tentative subdivision is propos-ed for the Uquian into three subages based on knowledge of the Mar del Plata-Miramar sequence, from oldest to youngest, the Barrancalobian, Vorohuean, and Sanandresian. In Argentina the Uquian is presently marked by the first known record of Scelidodon, Hydrochoeropsis, Ctenomys, Canidae, Ursidae, Gomphotheriidae, Equidae, Tapiridae, Camelidae, Cervidae, and the last known record of Thylatheridium, Thylophorops, Dankomys, Eumysops, Pithanotomys, Eucoelophorus, Hegetotheriidae, Sparassocynidae, and Microtragulidae.

The Ensenadan Age is based on the fauna from the Ensenada Formation near the city of Ensenada, Buenos Aires Province. In Argentina the Ensenadan is marked by the first known record of Lomaphorus, Neothoracophorus, Plaxhaplous, Cavia, Lyncodon, Lutra, Galera, Smilodon, Dicotyles, Lama, Vicugna, the last known record of Orthomyctera, and the only known record of Brachynasua.

     Typícal beds of late Lujanian Age in Argentina consist of fluvial deposits occupying stream channels, and shallow basins, often incised into beds of early Lujanian (i.e. Bonaerian of early workers) and Ensenadan Age. The Lujanian Age is based on a fauna from beds along the Rio Luján, about 65 km west of the city of Buenos Aires, Buenos Aires Province. The Lujanian in Argentina is marked by the first record of Equus, Chlamyphorus, and Holochilus, and the last record of Megatherioidea, Glyptodontoidea, Arctodus (=Arctotherium), Smilodon, Litopterna, Notoungulata, Proboscidea, Equidae, Morenelaphus, and Palaeolama.

   These land mammal ages are often difficult to recognize in other South American countries. The compositions of South American Pleistocene faunas vary with the environment. Some taxa were widely distributed in fossil deposits throughout the continent, but their occurrences need not reflect synchroneity. This is a result of changing climates and habitats in time. Consequently, proposed intracontinental correlations need confirmation based on magnetostratigraphy and a radioisotope time scale. Paleontologic characterizations of these land mammal ages (i.e. first and last record, and guide fossils) are useful for much of Argentina, but extensions to most of the other parts of South America are at best tenuous.

The majority of known non-Argentine Pleistocene faunas are believed to be Lujanian in age. Possible non Argentine early Pleistocene (Uquian) faunas include Ayo Ayo and Anzaldo in Bolivia, and Cocha Verde in southern Columbia. A possible middle Pleistocene (Ensenadan or early Lujanian) fauna is the Chichense of Ecuador. Paleomagnetic and radioisotopic date (MacFadden et al., 1983) clearly indicate that the greater part of the Tarija fauna (Bolivia) is Ensenadan in age.

  The end of the Pleistocene and beginning of the Holocene in South America is marked by extinction of nearly all large mammalian herbivores and their specialized large predators. Radiocarbon age determinations suggest that large scale extinctions of megafauna occurred between 15,000 and 8,000 yrs. B.P. (years before present). 

A new carcharhinid shark species, Physogaleus hemmooriensis sp. nov., is described from the Lower Hemmoorian (Behrendorfian, late Burdigalian, early Miocene) of Werder, Lower Saxony, Germany. P. hemmooriensis also occurs in the Edegem and Antwerpen Sands Members of the Berchem Formation, Belgium, and in the Miste Bed, Aalten Member of the Breda Formation, The Netherlands, which have an early to middle Miocene age. In the Western Atlantic region, the taxon is present in the early Miocene Calvert Formation of Delaware, U.S.A, which is largely contemporaneous with the Hemmoorian. 

A rear skull from the Quercy Phosphorites is described. It documents a new perissodactyl genus, likely assignable to the family Palaeotheriidae and probably paleogene of age. Owing to the lack of any tooth, this family assignment remains however somewhat hypothetical. The specimen is firstly characterised by the presence of a wide epitympanic sinus swelling and hollowing the squamosal shell and the post-glenoid process. This cavity might make up a peculiar pattern of improvement for the hearing apparatus by carrying out a kind of drum near the middle ear, whereas the bony tympanic remains barely bulged and forms but a few developed auditory bulla. This pattern appears an outcome of a trend observed with many palaeotheriids, such as Plagiolophus. Furthermore, the endocranial cast shows a rather high degree of gyrencephaly for a paleogene mammal. The prominent lambdoidal crest points out a powerful nape musculature and a lowered head bearing. Consequently, it is assumed that such an animal was probably living in somewhat open places, at the expense of rather tough vegetables. It might have been accordingly provided with a semi-hypsodont, cement covered, "plagiolophoid" dentition. 

Several endemic deer remains from the Middle Pleistocene deposits of the Castiglione cave (Oletta, Haute-Corse) are examined here. A morphometric analysis allows to relate them to a new insular subspecies Cervus elaphus rossii. The bones were compared with those of the mainland early Middle Pleistocene subspecies Cervus elaphus acoronatus Beninde and the European species Cervus elaphus Linné (Late Middle Pleistocene and Upper Pleistocene forms (continental and insular)). The Castiglione fossil shows peculiar morphofunctional features in its appendicular skeleton suggesting a morphological convergence with certain Bovidae. 

Le matériel provenant de nouvelles fouilles dans les phosphorites du Quercy, soumis aux divers spécialistes, a conduit à la soutenance de plusieurs thèses d'état: sur les rongeurs (Hartenberger, Vianey-Liaud), les Chiroptères et Insectivores (Sigé), les Artiodactyles (Sudre), les Squamates (Rage) et, en partie, les Chéloniens (De Broin). Chacun dans son domaine, à côté des conclusions d'ordre évolutif, paléogéographique ou paléoécologique, a pu établir des successions stratigraphiques des gisements du Quercy qui se sont révélées largement concordantes. Ainsi la succession des faunes du Quercy est actuellement l'une des mieux établies. Dans ce contexte, les différents spécialistes ont décidé de conjuguer leurs efforts dans l'étude monographique de plusieurs gisements repérés le long de cette échelle, afin de rassembler l'information paléontologique sur des faunes bien précises et de tenter d'obtenir des indications d'ordre taphonomique. 

Extensive collecting in previously known and new Middle Eocene rodent localities in the Subathu Group of the Rajauri and Reasi districts, Jammu and Kashmir (northwest Himalaya, India) has yielded over 500 isolated cheek teeth, numerous incisors and cheek tooth fragments, and two mandibular fragments with M/1-M/3. An analysis of this additional material and a restudy of that reported earlier from the Metka (Rajauri) area has revealed the presence of diverse Middle Eocene ctenodactyloid rodents in India. The assemblage comprises six genera and twelve species, viz.., Bírbalomys woodi, B. ibrahimshahi, B. sondaari, Basalomys vandermeuleni, B. ijlsti, B. lavocati, Chapattimys wilsoni, C. debruijni, Gumbatomys asifi, cf. Advenimus bohlini, cf. Petrokoslovia sp. indet. 1 and cf. Petrokoslovia sp. indet. 2. Most species of this assemblage are endemic to the Indian subcontinent and are referred to the Family Chapattimyidae. However, at least three taxa, viz.., cf. Advenimus bohlini, cf. Petrokoslovia sp.indet. 1 and cf. Pezrokoslovia sp. indet. 2 show close affinities with their Central and Eastern Asiatic contemporaries and are attributed to the Family Yuomyidae. The dentitions of most of the Middle Eocene rodents from India are described here in better detail thus providing more precise characterization than was available earlier. The taxonomie status of Basalomys, formerly a subgenus of Birbalomys has been raised to that of a genus. Basalomys vandermeuleni is here designated as the type species of Basalomys in place of B. ijlsti. The rodent assemblage from the Subathu Group is specifically similar to that from the coeval Kuldana Formation of Pakistan. The absence of Birbalomys ibrahimshahi and cf. Advenimus bohlini in Pakistan is related to sampling bias. The identifications of the Early Eocene rodents from Barbara Banda, Pakistan have been reexamined and the familial assignments of some of the Early and Middle Eocene rodents from Central and Eastern Asia have been discussed and reviewed in the light of recent discovery of early rodents in Mongolia.